Two thousand years ago one of history's most notorious individuals was born. Professor Mary Beard embarks on an investigative journey to explore the life and times of Gaius Julius Caesar Augustus Germanicus - better known to us as Caligula. Caligula has now become known as Rome's most capricious tyrant, and the stories told about him are some of the most extraordinary told about any Roman emperor. He was said to have made his horse a consul, proclaimed himself a living God, and indulged in scandalous orgies - even with his own three sisters - and that's before you mention building vast bridges across land and sea, prostituting senators' wives and killing half the Roman elite seemingly on a whim. All that in just four short years in power before a violent and speedy assassination in a back alley of his own palace at just 28-years-old." But how much of his story is true? Travelling across the Roman world - from Germany and Capri in the bay of Naples to the astonishing luxury of his life in imperial Rome - Mary attempts to peel away the myths. Some stories are difficult to get to the bottom of as they were written by hacks long after his death, but there is plenty of surviving evidence where the 'real' Caligula can be glimpsed. Such as in the extraordinary luxury of his private yachts outside Rome; in the designs he chose for his coins when he became Emperor; in an eye-witness account of Caligula's withering humour written in 41AD; the trial documents covering the mysterious death of his father when he was just seven-years-old; and even in a record of his imperial slaves - from the palace spy to his personal trainer. Piecing together the evidence, Mary puts Caligula back into the context of his times to reveal an astonishing story of murder, intrigue and dynastic family power. Above all, she explains why Caligula has ended up with such a seemingly unredeemable reputation. In the process, she reveals a more intriguing portrait of not just the monster, but the man.

Earthflight departs on its grand European tour, using a host of techniques including taking extraordinary footage from microlites as they fly alongside imprinted birds. Among other wonders, cameras soar with cranes and geese over Venice, the white cliffs of Dover and Edinburgh. White storks leave Africa and struggle to reach Istanbul, the gateway to Europe, while cranes take an easier route over the monkey-guarded Rock of Gibraltar. In France, the white horses of the Camargue make a splash around the new arrivals. In Rome, the heat of the city lures 20 million starlings that perform nature's greatest aerial display to outwit peregrines. In Hungary, sand martins grab mayflies from the air; and at Bass Rock, gannets create Britain's greatest natural spectacle as they dive in exquisite slow-motion. The story takes an emotional turn as storks and swallows wait for their partners to return and indulge in a spot of DIY to impress. Finally, geese touch down in Svalbard to raise a family. To protect their young, a squadron of birds assemble to see off polar bears. With views of birds flying over the Loire Valley, London docklands and the bulb fieds of Holland, this is Europe as never before.

Horses. Powerful, graceful, and thunderously fast. No animal has made a greater impact on society or given humans more freedom and mobility. In a spectacular adventure, the film unlocks the mystery of the first horse warriors on the vast, grassy plains of Kazakhstan, where horses still roam free, and nomadic herders follow their traditional way of life.
Investigating clues from archaeology and genetics, researchers reveal vivid evidence of the very first horsemen. They also discover warriors who swept across Europe and turn out to be the ancestors of millions today.

The third episode is devoted to the ways in which plants reproduce. Pollen and a stigma are the two components needed for fertilisation. Most plants carry both these within their flowers and rely on animals to transport the pollen from one to the stigma of another. To do this, they attract their couriers with colour, scent and nectar. It isn't just birds that help pollination: some mammals and reptiles also do so. However, it is mostly insects that are recruited to carry out the task. To ensure that pollen is not wasted by being delivered to the wrong flower, some species of plant have developed exclusive relationships with their visitors, and the gentian and its attendant carpenter bees is one example. Since pollen can be expensive to produce in terms of calories, some plants, such as orchids, ration it by means of pollinia and a strategically placed landing platform. Other orchids offer no reward for pollination, but instead mislead their guests by mimicking their markings and aroma, thus enticing males to 'mate' with them (Pseudocopulation). The most extreme fertilisation method is one of imprisonment, and one plant that uses it is the dead horse arum. It is often found near gull colonies, and mimics the appearance and smell of rotting flesh. Blow-flies are attracted to it, and are forced to stay the night before being allowed to depart in the morning, laden with pollen. Finally, Attenborough introduces the world's largest inflorescence: that of the titan arum.

A fifth of the land on our planet is covered by desert. Deserts may appear to be barren and empty, but they are of crucial importance to life. For those that can overcome their challenges they provide a vital refuge. The Grasslands are one of our planet's most productive landscapes. They support the greatest aggregations of large animals on Earth. Cameras follow desert elephants seeking sustenance, bison roaming North American grasslands and caterpillars living the good life underground.

One extraordinary family of fish has achieved the ultimate form of gender equality. When it comes to breeding, seahorses do it like no other animal on the planet. Their fierce females refuse to be left holding the baby. So it's the males that carry their brood. In this episode we explore the world of mating seahorses which are the only animals in the world that have a major gender role reversal.