Fourth episode examines how plants either share environments harmoniously or compete for dominance within them. Attenborough highlights the 1987 hurricane and the devastation it caused. However, for some species, it was that opportunity for which they had lain dormant for many years. The space left by uprooted trees is soon filled by others who move relatively swiftly towards the light. The oak is one of the strongest and longest-lived, and other, lesser plants nearby must wait until the spring to flourish before the light above is extinguished by leaves. Tropical forests are green throughout the year, so brute force is needed for a successful climb to the top of the canopy: the rattan is an example that has the longest stem of any plant. As its name suggests, the strangler fig 'throttles' its host by growing around it and cutting off essential water and light. Some can take advantage of a fallen tree by setting down roots on the now horizontal trunk and getting nutriment from the surrounding moss and the fungi on the dead bark. The mountain ash (eucalyptus regnans) grows so tall, that regeneration becomes a considerable problem. It is easily flammable, so its solution is to shed its seeds during a forest fire and sacrifice itself. It therefore relies on the periodic near-destruction of its surroundings in order to survive. Attenborough observes that catastrophes such as fire and drought, while initially detrimental to wildlife, eventually allow for deserted habitats to be reborn.

Mountain biking requires endurance, core strength and balance, bike handling skills, and self-reliance. Advanced riders pursue both steep technical descents and high incline climbs. In the case of freeriding, down hilling, and dirt jumping, aerial manoeuvres are performed off both natural features and specially constructed jumps and ramps. Follows several individuals breaking out of their mundane lives and entering into the adventurous thrill-seeking world of this sport.

Documentary telling the little-known story of how Darwin came to write his great masterpiece, On the Origin of Species, a book which explains the wonderful variety of the natural world as emerging out of death and the struggle of life. In the twenty years he took to develop a brilliant idea into a revolutionary book, Darwin went through a personal struggle every bit as turbulent as that of the natural world he observed. Fortunately, he left us an extraordinary record of his brilliant insights, observations of nature, and touching expressions of love and affection for those around him. He also wrote frank accounts of family tragedies, physical illnesses and moments of self-doubt, as he laboured towards publication of the book that would change the way we see the world.

Andrew Graham-Dixon travels to Istanbul to immerse himself in the tumultuous world of the Byzantine Empire. He decodes the iconography of the art of the period and explains its continuing relevance.

The very heart of Africa is covered in dense tropical rainforest. The animals that live here find the most ingenious ways to carve out their space in a claustrophobic landscape. Danger lurks in every shadow, but some animals thrive here, from honey-stealing chimps to birds with a lineage as old as the dinosaurs, thundering elephants and kick-boxing frogs. Here in the Congo, no matter how tough the competition, you must stand up and fight for yourself and your patch.

Tim Whewell is in St Petersburg to meet the self-styled Russian nationalists and patriots who are volunteering to join the fighting in eastern Ukraine. He joins a group of volunteers as they undertake military training, and travels to Ukraine to see how the volunteer force is fighting on the ground.