Our journey begins with David Attenborough at the North Pole, as the sun returns after six months of darkness. We follow a pair of courting polar bears, which reveal a surprisingly tender side. Next stop is the giant Greenland ice cap, where waterfalls plunge into the heart of the ice and a colossal iceberg carves into the sea. Humpback whales join the largest gathering of seabirds on earth to feast in rich Alaskan waters. Further south, the tree line marks the start of the Taiga forest, containing one third of all trees on earth. Here, 25 of the world's largest wolves take on formidable bison prey. At the other end of our planet, the Antarctic begins in the Southern Ocean where surfing penguins struggle to escape a hungry sea-lion and teams of orcas create giant waves to wash seals from ice floes -a filming first. Diving below the ice, we discover prehistoric giants, including terrifying sea spiders and woodlice the size of dinner plates. Above ground, crystal caverns ring the summit of Erebus, the most southerly volcano on earth. From here we retrace the routes of early explorers across the formidable Antarctic ice-cap - the largest expanse of ice on our planet. Finally, we rejoin David at the South Pole, exactly one hundred years after Amundsen then Scott were the first humans to stand there

There is no greater test for life than winter, as temperatures plummet to 70 below and winds reach 200kph. Darkness and ice extend across the polar regions and only a few remarkable survivors gamble on remaining. We join a female polar bear trekking into the Arctic mountains to give birth as the first blizzards arrive. Out on the frozen ocean, the entire world's population of spectacled eider ducks brave the winter in a giant ice hole kept open by ferocious currents. Arctic forests transform into a wonderland of frost and snow - the scene of a desperate and bloody battle between wolf and bison, but also where a remarkable alliance between raven and wolverine is made. Beneath the snow lies a magical world of winter survivors. Here tiny voles dodge the clutches of the great grey owl, but cannot escape the ultimate under-show predator - the least weasel. Midwinter and a male polar bear wanders alone across the dark, empty icescape. Below the snow, polar bear cubs begin life in an icy den while fantastical auroras light the night skies above. In Antarctica, we join male emperor penguins in their darkest hour, battling to protect precious eggs from fierce polar storms. Weddell seals escape to a hidden world of jewel-coloured corals and alien-looking creatures but frozen devastation follows as sinister ice stalactites reach down with deadly effect.

Mammals' ability to learn new tricks is the key to survival in the knife-edge world of hunters and hunted. In a TV first, a killer whale off the Falklands does something unique: it sneaks into a pool where elephant seal pups learn to swim and snatches them, saving itself the trouble of hunting in the open sea. Slow-motion cameras reveal the star-nosed mole's newly-discovered technique for smelling prey underwater: it exhales then inhales a bubble of air ten times per second. Young ibex soon learn the only way to escape a fox - run up an almost vertical cliff face - and young stoats fight mock battles, learning the skills that make them one of the world's most efficient predators.

For the animals in the polar regions, autumn means dramatic battles and epic journeys. Time is running out - the Arctic Ocean is freezing over and the sea ice is advancing at 2.5 miles per day around Antarctica. Polar bears gather in large numbers on the Arctic coast as they wait for the return of the ice. Soon, tempers fray and violent sparring contests break out. Meanwhile 2,000 beluga whales head for one special estuary, a gigantic 'whale spa' where they will thrash their snow-white bodies against the gravel and exfoliate. Inland, the tundra undergoes a dramatic transformation from green to fiery red. Here, musk ox males slam head-first into each other with the force of a 30mph car crash as they struggle to defend their harems. Frisky young caribou males play a game of 'grandma's footsteps' as they try to steal the boss's female. Down in Antarctica, Adelie penguin chicks huddle together in creches. When a parent returns from fishing, it leads its twins on a comical steeplechase - sadly there's only enough for one, so the winner gets the meal. Two months later and the chicks are fully feathered apart from downy Mohican hairdos - they're ready to take their first swim - reluctantly though, as it seems penguins are not born with a love of water! And with good reason - a leopard seal explodes from the sea and pulls one from an ice floe, a hunting manoeuvre that has never been filmed before. As winter approaches and everyone has left, the giant emperor penguin arrives and makes an epic trek inland to breed. The mothers soon return to the sea leaving the fathers to hold the eggs and endure the coldest winter on earth.

As almost all animal inhabitants of Antarctica are forced to migrate north, the sea underneath the frozen ice still provides a home to many specially adapted fish whose cells are protected from freezing through an 'antifreeze' liquid. Many of them feed on the faeces of other animals. The most notable larger animal that does not migrate north is perhaps the Weddell Seal, which can be found as close as 1300 kilometres to the pole. Groups of seals tear holes into the ice to dive for food and come up to breathe. The females come back to the ice to give birth. This episode also describes primitive plant life such as lichen, which can still be found on the continent in winter, even in the extremely dry and permanently frozen valleys conditions under which dead animals can lie frozen for many centuries without decomposing. It details the life of the Emperor Penguin, 'the only birds to lay their eggs directly on ice'. While other animals retreat, Emperors migrate not just to the ice, but into the Antarctic continent. The females lay eggs which are incubated by the males under the harshest conditions on Earth (huddling closely together for warmth), while the females return to the sea.

In the Land of the Cave Bear, Alice ventures to the parts of the northern hemisphere, hit hardest by the cold - Europe and Siberia. High in the mountains of Transylvania, a cave sealed for thousands of years reveals grisly evidence for a fight to the death between two staving giants, a cave bear and a cave lion. Yet Alice discovers that for woolly rhinos and woolly mammoths, the Ice Age created a bounty. The Mammoth Steppe, a vast tract of land which went half way round the world, provided food all year round, for those that liked the cold. It was these mammoths that Europe's most dangerous predators hunted for their survival.