Fresh water is our most precious resource and it defines the distribution of life on land. Follow the descent of rivers from their mountain sources to the sea. Watch spectacular waterfalls, fly inside the Grand Canyon and explore the wildlife below the ice in the world's deepest lake. Witness unique and dramatic moments of animal behaviour: a showdown between smooth-coated otters and mugger crocodiles; deep-diving long tailed macaques; massive flocks of snow geese on the wing and a piranha frenzy in the perilous waters of the world's largest wetland.

The Cave of Swallows in Mexico is a 400m vertical shaft, deep enough to engulf the Empire State Building. The Lechuguilla cave system in the USA is 193km long and 500m deep with astonishing crystal formations hanging from its chambers. Although often overlooked, caves are remarkable habitats with equally bizarre wildlife. Cave angel fish cling to the walls behind cave waterfalls with microscopic hooks on their flattened fins. Cave swiftlets navigate by echo-location and build nests out of saliva. The Texas cave salamander has neither eyes nor pigment. Unique access to a hidden world of stalactites, stalagmites, snotites and troglodytes brings a wealth of surprises.

Around 30% of the land's surface is desert, the most varied of our ecosystems despite the lack of rain. Unravel the secrets of desert survival and experience the ephemeral nature of this dynamic environment. Watch Saharan sandstorms nearly a mile high and desert rivers that run for a single day. In the Gobi Desert, rare Bactrian camels get moisture from the snow. In the Atacama, guanacos survive by licking dew off cactus spines. In the USA, the brief blooming of Death Valley triggers a plague of locusts 65km wide and 160km long. A unique aerial voyage over the Namibian desert reveals elephants on a long trek for food and desert lions searching for wandering oryx.

Jungles cover roughly three per cent of our planet yet contain 50 per cent of the world's species. High-definition cameras enable unprecedented views of animals living on the dark jungle floor. In the Ngogo forest the largest chimpanzee group in the world defends its territory from neighbouring groups. Other jungle specialists include parasitic fungi which infiltrate an insect host, feed on it, and then burst out of its body.

A humpback whale mother and calf embark on an epic journey from tropical coral paradises to storm ravaged polar seas. Newly discovered coral reefs in Indonesia reveal head-butting pygmy seahorses, flashing 'electric' clams and bands of sea kraits, 30-strong, which hunt in packs. Elsewhere plagues of sea urchins fell forests of giant kelp. Huge bull fur seals attack king penguins, who despite their weight disadvantage, put up a spirited defence.

The Taiga forest, on the edge of the Arctic, is a silent world of stunted conifers. The trees may be small but filming from the air reveals its true scale. A third of all trees on Earth grow here and during the short summer they produce enough oxygen to change the atmosphere. In California General Sherman, a giant sequoia, is the largest living thing on the planet, ten times the size of a blue whale. The oldest organisms alive are bristlecone pines. At more than 4,000 years old they pre-date the pyramids. But the baobab forests of Madagascar are perhaps the strangest of all.