This is the story of two tiny animals coming of age. In the wild woods of North America, a young chipmunk is gathering a vital store of nuts ahead of his first winter - in his way are ruthless rivals and giant predators. In the steaming rainforest, a young tree-shrew is forced deep into the jungle to find food. She must draw on all her intelligence and agility if she is to escape the ultimate jungle predator - a reticulated python!

About 340 million years ago a brand new family of animals was evolving in the primeval swamps. They were to go one step further than the amphibians who had emerged onto dry and before them. For they would eventually completely cut their ties with water. They were the ancestors of todays lizards. They evolved scaly impermeable skins and moved up into the forests. They diversified into a multitude of different shapes and sizes. They developed signalling systems to communicate with one another. And they squabbled as animals do. For food they hunted insects that were already well established on the land in great numbers. And here without returning to water they produced their families. They powered their bodies not only with food but with the heat that they drew directly from the sun.
As they diversified so they spread into the harshest of the lands habitats. The baking waterless deserts which eventually they would come to dominate. Discover jacky lizards that wave, wrestling beaded lizards and the the world's smallest chameleon, which is no bigger than his thumbnail, and the biggest lizard in Australia.

Spring arrives in the polar regions, and the sun appears after an absence of five months; warmth and life return to these magical ice worlds - the greatest seasonal transformation on our planet is underway. Male Adelie penguins arrive in Antarctica to build their nests - it takes a good property to attract the best mates and the males will stop at nothing to better their rivals! But these early birds face the fiercest storms on the planet. In the Arctic, a polar bear mother is hunting with her cubs. Inland, the frozen rivers start to break up and billions of tons of ice are swept downstream in the greatest of polar spectacles. This melt-water fertilizes the Arctic Ocean, feeding vast shoals of Arctic cod and narwhal. The influx of freshwater accelerates the breakup of the sea-ice - an area of ice the size of Australia will soon vanish from the Arctic. On land, a woolly bear caterpillar emerges from the snow having spent the winter frozen solid. Caterpillars normally become moths within months of hatching, but life is so harsh here that the woolly bear takes 14 years to reach adulthood. Once mature it has only days to find a mate before it dies! Alongside the caterpillars white Arctic wolves race to raise their adorable cubs before the cold returns.

It is high summer in the Polar Regions, and the sun never sets. Vast hordes of summer visitors cram a lifetime of drama into one long, magical day; they must feed, fight and rear their young in this brief window of plenty. Summer is a tough time for the polar bear family, as their ice world melts away and the cubs take their first swimming lesson. Some bears save energy by dozing on icy sun beds, while others go egg-collecting in an Arctic tern colony, braving bombardment by sharp beaks. There are even bigger battles on the tundra; a herd of musk oxen gallop to the rescue as a calf is caught in a life and death struggle with a pair of Arctic wolves. But summer also brings surprises, as a huge colony of 400,000 king penguins cope with an unlikely problem - heat. The adults go surfing, while the woolly-coated chicks take a cooling mud bath. Nearby, a bull fur seal is prepared to fight to the death with a rival. Fur flies as the little pups struggle desperately to keep out of the way of the duelling giants. Further south, a minke whale is hunted amongst the ice floes by a family of killer whales. The dramatic chase lasts over 2 hours and has never been filmed before. The killers harry the minke whale, taking it in turns to wear it down. Eventually it succumbs to the relentless battering. Finally, comical adelie penguins waddle back to their half a million strong colony like clockwork toys. The fluffy chicks need constant feeding and protection as piratical skuas patrol the skies. When an unguarded chick is snatched, a dramatic "dogfight" ensues.

For the animals in the polar regions, autumn means dramatic battles and epic journeys. Time is running out - the Arctic Ocean is freezing over and the sea ice is advancing at 2.5 miles per day around Antarctica. Polar bears gather in large numbers on the Arctic coast as they wait for the return of the ice. Soon, tempers fray and violent sparring contests break out. Meanwhile 2,000 beluga whales head for one special estuary, a gigantic 'whale spa' where they will thrash their snow-white bodies against the gravel and exfoliate. Inland, the tundra undergoes a dramatic transformation from green to fiery red. Here, musk ox males slam head-first into each other with the force of a 30mph car crash as they struggle to defend their harems. Frisky young caribou males play a game of 'grandma's footsteps' as they try to steal the boss's female. Down in Antarctica, Adelie penguin chicks huddle together in creches. When a parent returns from fishing, it leads its twins on a comical steeplechase - sadly there's only enough for one, so the winner gets the meal. Two months later and the chicks are fully feathered apart from downy Mohican hairdos - they're ready to take their first swim - reluctantly though, as it seems penguins are not born with a love of water! And with good reason - a leopard seal explodes from the sea and pulls one from an ice floe, a hunting manoeuvre that has never been filmed before. As winter approaches and everyone has left, the giant emperor penguin arrives and makes an epic trek inland to breed. The mothers soon return to the sea leaving the fathers to hold the eggs and endure the coldest winter on earth.

There is no greater test for life than winter, as temperatures plummet to 70 below and winds reach 200kph. Darkness and ice extend across the polar regions and only a few remarkable survivors gamble on remaining. We join a female polar bear trekking into the Arctic mountains to give birth as the first blizzards arrive. Out on the frozen ocean, the entire world's population of spectacled eider ducks brave the winter in a giant ice hole kept open by ferocious currents. Arctic forests transform into a wonderland of frost and snow - the scene of a desperate and bloody battle between wolf and bison, but also where a remarkable alliance between raven and wolverine is made. Beneath the snow lies a magical world of winter survivors. Here tiny voles dodge the clutches of the great grey owl, but cannot escape the ultimate under-show predator - the least weasel. Midwinter and a male polar bear wanders alone across the dark, empty icescape. Below the snow, polar bear cubs begin life in an icy den while fantastical auroras light the night skies above. In Antarctica, we join male emperor penguins in their darkest hour, battling to protect precious eggs from fierce polar storms. Weddell seals escape to a hidden world of jewel-coloured corals and alien-looking creatures but frozen devastation follows as sinister ice stalactites reach down with deadly effect.