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It is high summer in the Polar Regions, and the sun never sets. Vast hordes of summer visitors cram a lifetime of drama into one long, magical day; they must feed, fight and rear their young in this brief window of plenty. Summer is a tough time for the polar bear family, as their ice world melts away and the cubs take their first swimming lesson. Some bears save energy by dozing on icy sun beds, while others go egg-collecting in an Arctic tern colony, braving bombardment by sharp beaks. There are even bigger battles on the tundra; a herd of musk oxen gallop to the rescue as a calf is caught in a life and death struggle with a pair of Arctic wolves. But summer also brings surprises, as a huge colony of 400,000 king penguins cope with an unlikely problem - heat. The adults go surfing, while the woolly-coated chicks take a cooling mud bath. Nearby, a bull fur seal is prepared to fight to the death with a rival. Fur flies as the little pups struggle desperately to keep out of the way of the duelling giants. Further south, a minke whale is hunted amongst the ice floes by a family of killer whales. The dramatic chase lasts over 2 hours and has never been filmed before. The killers harry the minke whale, taking it in turns to wear it down. Eventually it succumbs to the relentless battering. Finally, comical adelie penguins waddle back to their half a million strong colony like clockwork toys. The fluffy chicks need constant feeding and protection as piratical skuas patrol the skies. When an unguarded chick is snatched, a dramatic "dogfight" ensues.

The documentary series reveals the extraordinary riches and wonders of the Polar Regions that have kept people visiting them for thousands of years. Today, their survival relies on a combination of ancient wisdom and cutting-edge science. Most Arctic people live in Siberia, either in cities like Norilsk - the coldest city on earth - or out on the tundra, where tribes like the Dogan survive by herding reindeer, using them to drag their homes behind them. On the coast, traditional people still hunt walrus from open boats - it is dangerous work, but one big walrus will feed a family for weeks. Settlers are drawn to the Arctic by its abundant minerals; the Danish Armed Forces maintain their claim to Greenland's mineral wealth with an epic dog sled patrol, covering 2,000 miles through the winter. Above, the spectacular northern lights can disrupt power supplies so scientists monitor it constantly, firing rockets into it to release a cloud of glowing smoke 100 kilometres high. In contrast, Antarctica is so remote and cold that it was only a century ago that the first people explored the continent. Captain Scott's hut still stands as a memorial to these men. Science is now the only significant human activity allowed; robot submarines are sent deep beneath the ice in search of new life-forms, which may also be found in a labyrinth of ice caves high up on an active volcano. Above, colossal balloons are launched into the purest air on earth to detect cosmic rays. At the South Pole there is a research base designed to withstand the world's most extreme winters. Cut off from the outside world for six months, the base is totally self-sufficient, even boasting a greenhouse.

Our journey begins with David Attenborough at the North Pole, as the sun returns after six months of darkness. We follow a pair of courting polar bears, which reveal a surprisingly tender side. Next stop is the giant Greenland ice cap, where waterfalls plunge into the heart of the ice and a colossal iceberg carves into the sea. Humpback whales join the largest gathering of seabirds on earth to feast in rich Alaskan waters. Further south, the tree line marks the start of the Taiga forest, containing one third of all trees on earth. Here, 25 of the world's largest wolves take on formidable bison prey. At the other end of our planet, the Antarctic begins in the Southern Ocean where surfing penguins struggle to escape a hungry sea-lion and teams of orcas create giant waves to wash seals from ice floes -a filming first. Diving below the ice, we discover prehistoric giants, including terrifying sea spiders and woodlice the size of dinner plates. Above ground, crystal caverns ring the summit of Erebus, the most southerly volcano on earth. From here we retrace the routes of early explorers across the formidable Antarctic ice-cap - the largest expanse of ice on our planet. Finally, we rejoin David at the South Pole, exactly one hundred years after Amundsen then Scott were the first humans to stand there

There is no greater test for life than winter, as temperatures plummet to 70 below and winds reach 200kph. Darkness and ice extend across the polar regions and only a few remarkable survivors gamble on remaining. We join a female polar bear trekking into the Arctic mountains to give birth as the first blizzards arrive. Out on the frozen ocean, the entire world's population of spectacled eider ducks brave the winter in a giant ice hole kept open by ferocious currents. Arctic forests transform into a wonderland of frost and snow - the scene of a desperate and bloody battle between wolf and bison, but also where a remarkable alliance between raven and wolverine is made. Beneath the snow lies a magical world of winter survivors. Here tiny voles dodge the clutches of the great grey owl, but cannot escape the ultimate under-show predator - the least weasel. Midwinter and a male polar bear wanders alone across the dark, empty icescape. Below the snow, polar bear cubs begin life in an icy den while fantastical auroras light the night skies above. In Antarctica, we join male emperor penguins in their darkest hour, battling to protect precious eggs from fierce polar storms. Weddell seals escape to a hidden world of jewel-coloured corals and alien-looking creatures but frozen devastation follows as sinister ice stalactites reach down with deadly effect.

The polar regions of our planet may seem beyond the reach of most of us, but they are not beyond our influence. We, unintentionally, are changing these frozen worlds, and these changes will not just affect the poles but the whole planet. On the unforgiving frontier of climate change, polar bears, walruses, seals and penguins find their icy Edens in peril.

The episode is a journey through the human gut and the process of turning food into energy to keep the body going. Humans have turned eating into culture, a way to celebrate our roots, to bring us together. But food is really just one thing: Fuel. It's where you get the power to live your life. Even when you're sitting completely still, your body needs a lot of energy to stay on. But nowhere is the need for fuel more apparent than when you push yourself to the absolute limit. And the microbiome, the bacteria living in our gut, plays a great role in all this.