Caesar has been murdered, Mark Antony emerges from the Senate in shock, only to face Quintus Pompey and his thugs, who immediately follow him to try to kill him. Brutus returns home shaking after the murder. His mother is already plotting the return of the Roman Republic. Titus Pullo asks Eirene to marry him and she accepts. Erastes Fulmen kidnaps Lucius Vorenus' children and sister-in-law.
Mark Antony proposes an amnesty to the rest of the senate to keep the peace and allow the Republic to continue. Vorenus says goodbye to Niobe while Rome says goodbye to Caesar. Vorenus and Pullo track down Fulmen to a bath house, where he tells them he killed Vorenus' family.

As they prepare their counterattack against Caesar, Pompey's Republicans begin to squabble over both the anticipated spoils and the method of finishing Caesar off. Badly outnumbered, Caesar and Antony make their stand at Pharsalus. As they try to reach Caesar in Greece, the shipwrecked Vorenus and Pullo confront an unexpected dilemma.
Back in Rome, Atia again sends Octavia to beg protection from Servilia, who complies readily to encourage her growing friendship with Octavia. Niobe's sister Lyde comes back to see her and comforts her over Vorenus's disappearance.

Cicero is finally undone by his duplicity in the form of Titus Pullo, acting upon Octavian's request to assassinate all supporters of Brutus in Rome. Pullo and Vorenus enjoy a happy if counterfeit outing with their families in the countryside to hide their assassin mission.
Vorena the Elder's dangerous dalliance with one of Mimeo's men continues, and the torrid affair between Octavia and Agrippa escalates. Atia's casual cruelty to ask for the assassination of people she doesn't like results in complete disaster for Octavia's friend Jocasta.

Cities are growing at a faster rate than any other habitat on Earth. They may seem an unlikely place for animals to thrive, but they can be a world of surprising opportunity. Leopards prowl the streets of Mumbai, peregrine falcons hunt amongst New York's skyscrapers, and a million starlings perform spectacular aerial dances over Rome. In Jodhpur, langurs are revered as religious deities and in Harar, locals live in harmony with wild hyenas. Many animals, however, struggle to cope in the urban jungle. As the architects of this environment, can humans choose to build cities that are homes for both them and wildlife?

In the fourth episode, Dromaeosaurs stalk a hadrosaur herd as they cross a freezing river. Male Ornithomimus raid rival nests to bolster their own. Olorotitan raise their offspring on fertile volcanic fields but contend with biting mosquitoes.
Troodontids hunt mammals flushed out by a forest fire. A juvenile Antarctopelta scours the forest for a new winter den. Within the snow-covered forest, a tense standoff develops between ancient rivals, Pachyrhinosaurus and Nanuqsaurus.

Thanks to stunning reconstitutions with computer-generated images, this documentary retraces the manufacturing secrets of the largest and most exceptional city of Antiquity: Rome, the capital of the Mediterranean world. Three of its most emblematic monuments come to life. The Pantheon and its majestic dome, the sumptuous palace of Nero which covered a quarter of the city, and finally the legendary Colosseum, the largest Roman amphitheater ever built.