At the end of each Antarctic summer, the emperor penguins of the South Pole journey to their traditional breeding grounds in a fascinating mating ritual that is captured in this documentary by intrepid filmmaker Luc Jacquet. The journey across frozen tundra proves to be the simplest part of the ritual, as after the egg is hatched, the female must delicately transfer it to the male and make her way back to the distant sea to nourish herself and bring back food to her newborn chick.

Our journey begins with David Attenborough at the North Pole, as the sun returns after six months of darkness. We follow a pair of courting polar bears, which reveal a surprisingly tender side. Next stop is the giant Greenland ice cap, where waterfalls plunge into the heart of the ice and a colossal iceberg carves into the sea. Humpback whales join the largest gathering of seabirds on earth to feast in rich Alaskan waters. Further south, the tree line marks the start of the Taiga forest, containing one third of all trees on earth. Here, 25 of the world's largest wolves take on formidable bison prey. At the other end of our planet, the Antarctic begins in the Southern Ocean where surfing penguins struggle to escape a hungry sea-lion and teams of orcas create giant waves to wash seals from ice floes -a filming first. Diving below the ice, we discover prehistoric giants, including terrifying sea spiders and woodlice the size of dinner plates. Above ground, crystal caverns ring the summit of Erebus, the most southerly volcano on earth. From here we retrace the routes of early explorers across the formidable Antarctic ice-cap - the largest expanse of ice on our planet. Finally, we rejoin David at the South Pole, exactly one hundred years after Amundsen then Scott were the first humans to stand there

It is high summer in the Polar Regions, and the sun never sets. Vast hordes of summer visitors cram a lifetime of drama into one long, magical day; they must feed, fight and rear their young in this brief window of plenty. Summer is a tough time for the polar bear family, as their ice world melts away and the cubs take their first swimming lesson. Some bears save energy by dozing on icy sun beds, while others go egg-collecting in an Arctic tern colony, braving bombardment by sharp beaks. There are even bigger battles on the tundra; a herd of musk oxen gallop to the rescue as a calf is caught in a life and death struggle with a pair of Arctic wolves. But summer also brings surprises, as a huge colony of 400,000 king penguins cope with an unlikely problem - heat. The adults go surfing, while the woolly-coated chicks take a cooling mud bath. Nearby, a bull fur seal is prepared to fight to the death with a rival. Fur flies as the little pups struggle desperately to keep out of the way of the duelling giants. Further south, a minke whale is hunted amongst the ice floes by a family of killer whales. The dramatic chase lasts over 2 hours and has never been filmed before. The killers harry the minke whale, taking it in turns to wear it down. Eventually it succumbs to the relentless battering. Finally, comical adelie penguins waddle back to their half a million strong colony like clockwork toys. The fluffy chicks need constant feeding and protection as piratical skuas patrol the skies. When an unguarded chick is snatched, a dramatic "dogfight" ensues.

Mammals dominate the planet. They do it through having warm blood and by the care they lavish on their young. Weeks of filming in the bitter Antarctic winter reveal how a mother Weddell seal wears her teeth down keeping open a hole in the ice so she can catch fish for her pup. A powered hot air balloon produces stunning images of millions of migrating bats as they converge on fruiting trees in Zambia, and slow-motion cameras reveal how a mother rufous sengi exhausts a chasing lizard. A gyroscopically stabilised camera moves alongside migrating caribou, and a diving team swim among the planet's biggest fight as male humpback whales battle for a female.

There is no greater test for life than winter, as temperatures plummet to 70 below and winds reach 200kph. Darkness and ice extend across the polar regions and only a few remarkable survivors gamble on remaining. We join a female polar bear trekking into the Arctic mountains to give birth as the first blizzards arrive. Out on the frozen ocean, the entire world's population of spectacled eider ducks brave the winter in a giant ice hole kept open by ferocious currents. Arctic forests transform into a wonderland of frost and snow - the scene of a desperate and bloody battle between wolf and bison, but also where a remarkable alliance between raven and wolverine is made. Beneath the snow lies a magical world of winter survivors. Here tiny voles dodge the clutches of the great grey owl, but cannot escape the ultimate under-show predator - the least weasel. Midwinter and a male polar bear wanders alone across the dark, empty icescape. Below the snow, polar bear cubs begin life in an icy den while fantastical auroras light the night skies above. In Antarctica, we join male emperor penguins in their darkest hour, battling to protect precious eggs from fierce polar storms. Weddell seals escape to a hidden world of jewel-coloured corals and alien-looking creatures but frozen devastation follows as sinister ice stalactites reach down with deadly effect.

For the animals in the polar regions, autumn means dramatic battles and epic journeys. Time is running out - the Arctic Ocean is freezing over and the sea ice is advancing at 2.5 miles per day around Antarctica. Polar bears gather in large numbers on the Arctic coast as they wait for the return of the ice. Soon, tempers fray and violent sparring contests break out. Meanwhile 2,000 beluga whales head for one special estuary, a gigantic 'whale spa' where they will thrash their snow-white bodies against the gravel and exfoliate. Inland, the tundra undergoes a dramatic transformation from green to fiery red. Here, musk ox males slam head-first into each other with the force of a 30mph car crash as they struggle to defend their harems. Frisky young caribou males play a game of 'grandma's footsteps' as they try to steal the boss's female. Down in Antarctica, Adelie penguin chicks huddle together in creches. When a parent returns from fishing, it leads its twins on a comical steeplechase - sadly there's only enough for one, so the winner gets the meal. Two months later and the chicks are fully feathered apart from downy Mohican hairdos - they're ready to take their first swim - reluctantly though, as it seems penguins are not born with a love of water! And with good reason - a leopard seal explodes from the sea and pulls one from an ice floe, a hunting manoeuvre that has never been filmed before. As winter approaches and everyone has left, the giant emperor penguin arrives and makes an epic trek inland to breed. The mothers soon return to the sea leaving the fathers to hold the eggs and endure the coldest winter on earth.