The second programme deals with the mechanics of flight. Getting into the air is by far the most exhausting of a bird's activities, and Sir Attenborough observes shearwaters in Japan that have taken to climbing trees to give them a good jumping-off point. The albatross is so large that it can only launch itself after a run-up to create a flow of air over its wings. A combination of aerodynamics and upward air currents (or thermals), together with the act of flapping or gliding is what keeps a bird aloft. Landing requires less energy but a greater degree of skill, particularly for a big bird, such as a swan. Weight is kept to a minimum by having a beak made of keratin instead of bone, a light frame, and a coat of feathers, which is maintained fastidiously. The peregrine falcon holds the record for being fastest in the air, diving at speeds of over 300 km/h. Conversely, the barn owl owes its predatory success to flying slowly, while the kestrel spots its quarry by hovering. However, the true specialists in this regard are the hummingbirds, whose wings beat at the rate of 25 times a second. The habits of migratory birds are explored. After stocking up with food during the brief summer of the north, such species will set off on huge journeys southwards. Some, such as snow geese, travel continuously, using both the stars and the sun for navigation. They are contrasted with hawks and vultures, which glide overland on warm air, and therefore have to stop overnight.

Cities are growing at a faster rate than any other habitat on Earth. They may seem an unlikely place for animals to thrive, but they can be a world of surprising opportunity. Leopards prowl the streets of Mumbai, peregrine falcons hunt amongst New York's skyscrapers, and a million starlings perform spectacular aerial dances over Rome. In Jodhpur, langurs are revered as religious deities and in Harar, locals live in harmony with wild hyenas. Many animals, however, struggle to cope in the urban jungle. As the architects of this environment, can humans choose to build cities that are homes for both them and wildlife?

David Attenborough concludes his epic history of the evolution of flight with an exploration of the highly advanced fliers that dominate our skies today – the extraordinarily diverse skills of the birds, and the sonar-guided precision of the bats. He encounters some of our planet’s most remarkable fliers – from peregrine falcons dive-bombing starlings over Rome, to hummingbirds hovering in the cloud forests of Ecuador. In a spectacular finale, he visits Gomantong cave in Borneo to witness the mass exodus of a million bats.

This episode examines those birds whose sustenance comes from flesh and their methods of hunting. In New Zealand, Sir Attenborough observes Keas, parrots that do not eat meat exclusively, raiding a shearwater's burrow for a chick. However, it is the dedicated birds of prey, such as owls, buzzards, eagles, falcons and vultures, to which much of the programme is devoted. In order to spot and pursue their victims, senses of sight and hearing are very acute. Vultures are the exception, in that they eat what others have left, and once a carcass is found, so many birds descend on it that the carrion seems submerged beneath them. The Turkey Vulture is an anomaly within its group, as it also has a keen sense of smell. Eagles defend their territory vigorously, and a pair of sea eagles are shown engaging in an aerial battle. The Galápagos Hawk hunts Marine Iguanas, but can only do so when its quarry is vulnerable, during the breeding season. The African Harrier Hawk has adapted to extracting burrowing animals by virtue of an especially long, double-jointed pair of legs. By contrast, a shrike is not equipped with the requisite sharp beak and talons needed for butchery, and so dismembers its kill by impaling it on the thorns of acacias. The Lammergeier eats bones, and will drop them on to rocks from a great height in order to break them down to a digestible size. Also featured are the Eurasian Sparrowhawk, Goshawk and Peregrine Falcon.

Earthflight departs on its grand European tour, using a host of techniques including taking extraordinary footage from microlites as they fly alongside imprinted birds. Among other wonders, cameras soar with cranes and geese over Venice, the white cliffs of Dover and Edinburgh. White storks leave Africa and struggle to reach Istanbul, the gateway to Europe, while cranes take an easier route over the monkey-guarded Rock of Gibraltar. In France, the white horses of the Camargue make a splash around the new arrivals. In Rome, the heat of the city lures 20 million starlings that perform nature's greatest aerial display to outwit peregrines. In Hungary, sand martins grab mayflies from the air; and at Bass Rock, gannets create Britain's greatest natural spectacle as they dive in exquisite slow-motion. The story takes an emotional turn as storks and swallows wait for their partners to return and indulge in a spot of DIY to impress. Finally, geese touch down in Svalbard to raise a family. To protect their young, a squadron of birds assemble to see off polar bears. With views of birds flying over the Loire Valley, London docklands and the bulb fieds of Holland, this is Europe as never before.

Each night, across the planet, our cities illuminate the darkness. Their lights burning brighter than ever before. And whilst we're asleep, animals are making the most of this new neon landscape. The sunset in Los Angeles, Chicago, and Toronto creates worlds of opportunities for the cleverest animals willing to take the risk.