The final episode explores dinosaurs' extraordinary ability to survive. Featured dinosaurs include the bizarre magyarosaurus, which lived in the shadow of the biggest flying animal - hatzegopteryx - and showed an amazing adaptation to island life; and the weird nothronychus, a carnivore that gave up meat eating. This astonishing capacity to evolve into ever more diverse and bizarre forms meant that dinosaurs not only spread throughout the world, but also dominated life upon it for more than 160 million years. It was only an unprecedented extraterrestrial event that finally saw the end of planet dinosaur.

What triggered the incredible diversity of life on Earth, and how have complex life forms, including humans, evolved? Is there direction to evolution? And is human intelligence inevitable? This program focuses on evolution's 'great transformations' —among them the development of a standard four-limbed body plan, the journey from water to land, the return of marine mammals to the sea, and the emergence of humans. Driven by a combination of opportunism and a genetic 'toolkit', these astounding leaps forward define the arc of evolution. And they suggest that every living creature on earth today, and every species that has ever existed, is a variation on a grand genetic theme—a member of one, and only one, tree of life

This scientific detective tale tells the story of a remarkable group of pioneers who wanted to reach the ultimate extreme: absolute zero, a place so cold that the physical world as we know it doesn't exist, electricity flows without resistance, fluids defy gravity and the speed of light can be reduced to 38 miles per hour. Absolute zero became the Holy Grail of temperature physicists and is considered the gateway to many new technologies, such as nano-construction, neurological networks and quantum computing. The possibilities, it seems, are limitless. The first episode Chronicles the major discoveries leading towards the mastery of cold, beginning with King James I's court magician, Cornelius Drebbel, who managed to air condition the largest interior space in the British Isles in 1620. Other stories will include the first "natural philosopher," Robert Boyle, a founder of the Royal Society in Great Britain; the Grand Duke Ferdinand II de Medici's involvement in the creation of the first thermometer; the establishment of the laws of thermodynamics by three young scientists, Sadi Carnot, James Joule and William Thomson; and Michael Faraday's critical achievement in liquefying several other gases which set the stage for the commercial application of cold to refrigeration and air conditioning.

Silk is the invertebrates' great invention, used in a range of ways from from the protective stalks of lacewing eggs to the amazing hanging threads of New Zealand's 'glow worms'. Spiders, though, have taken silk-spinning to extremes. The common wolf spider has no web, but the female is a gentle parent, encasing her eggs in silk and carrying the precious bundle wherever she goes. The bolas spider uses a ball of sticky silk soaked in a copy of moth pheromone to lure its prey. Millions of communal spiders live and feed together in a vast, towering web - an arachnophobe's nightmare.

A visit to arguably the most famous archipelago on Earth, the Galapagos. It's home to a myriad of bizarre and unique creatures, endemic to these islands - but how did they get here and what is the key to these extraordinary islands that allows them to thrive? The programme reveals that this key holds not just the secret to life here, but also to how Darwin was able to leave with the ideas that would revolutionise biology.

Lucy Worsley continues her journey through Russia in the footsteps of the Romanovs, the most powerful royal dynasty in modern European history. In this episode she examines the extraordinary reign of Catherine the Great, and the traumatic conflict with napoleonic France that provides the setting for the novel War and Peace. At the magnificent palace of Peterhof near St Petersburg, Lucy charts the meteoric rise of Catherine the Great, who seized the Russian throne from her husband Peter III in 1762 and became the most powerful woman in the world. Catherine was a woman of huge passions - for art, for her adopted country (she was German by birth) and for her many lovers.
Catherine expanded her empire through military victories overseas, while at home she encouraged education and introduced smallpox inoculation to Russia. But Catherine struggled to introduce deeper reforms, and the institution of serfdom remained largely unchanged. Lucy explains how this injustice fuelled a violent rebellion. Nevertheless, Catherine left Russia more powerful on the world stage than ever. But all she had achieved looked set to be undone when Napoleon invaded Russia in 1812. Lucy relives the pivotal battle of Borodino, when the Russian army finally confronted the French forces; the traumatic destruction of Moscow; and, under Catherine's grandson Alexander, the eventual victory over the French that provided the Romanov dynasty with its most glorious hour.