This episode tells the story of how the Emperor Constantine brought Christianity to the western world. In AD 312, Rome was in crisis. The empire had been divided into four parts, each with its own emperor who fought one another. Constantine intervened and united Rome, using military might and a new religion - Christianity.

Deep in the Bolivian Andes at the height of 13,000ft stands Tiwanaku, the awe-inspiring ruins of a monolithic temple city. Built by a civilisation who dominated a vast swathe of South America, it was abandoned 1,000 years ago. For centuries it has been a mystery - how did a civilisation flourish at such an altitude and why did it vanish? Jago Cooper journeys through Bolivia's spectacular landscape to investigate the origins of Tiwanaku and finds evidence of an ancient people with amazing understanding of their environment, whose religion was based on collective effort and ritual beer drinking.

In the fourth episode of the series, Commodus defeats his sister's coup but withdraws from governing, allowing others to amass power while he engages in an affair and banishes his wife.

In Egypt, Caesar rebukes the advisers of the boy king, Ptolemy XIII, for their presumption in eliminating Pompey and demands the man who killed him. Caesar decides to intervene in the dispute between Ptolemy and his sister-wife, Cleopatra, to ensure both Rome's grain supply and his own access to Egypt's treasure.
Antony, in an unusual show of prudence, advises against this with only half a legion in Alexandria and Cato and Scipio still at large in Africa. In Rome, Brutus gets a chilly reception from Servilia when he returns home from Greece.

In AD 410, the Goth hordes sacked the city. This event symbolised Rome's collapse. Within 70 years, the western empire - what we think of as Ancient Rome - was abandoned. But it should never have happened at all.

The Bronze Age had the first large urban centers, powerful kingdoms and armies, writing, and trade routes across vast areas. What led to the collapse of the city-state entities and the end of the Bronze Age? Possibly a perfect storm of uncontrollable factors that are still being researched.