In Africa's ancient south west corner, two extraordinary deserts sit side by side. Water is in short supply, yet these deserts are somehow full of life because the creatures that live here have turned the rules of survival on their head. This film celebrates nature's ingenuity, no matter how tough it gets. In the Kalahari scrublands, clever meerkats are outsmarted by a wily bird, solitary and belligerent black rhinos get together to party and giant insects stalk huge flocks of birds. Rain almost never falls in the Namib - instead it must make do with vaporous, vanishing fog. The creatures in this, the world's oldest desert, have gone to the extremes, as spiders wheel to escape and a desert giraffe fights to defend his scant resources in the greatest giraffe battle ever filmed.

The Cave of Swallows in Mexico is a 400m vertical shaft, deep enough to engulf the Empire State Building. The Lechuguilla cave system in the USA is 193km long and 500m deep with astonishing crystal formations hanging from its chambers. Although often overlooked, caves are remarkable habitats with equally bizarre wildlife. Cave angel fish cling to the walls behind cave waterfalls with microscopic hooks on their flattened fins. Cave swiftlets navigate by echo-location and build nests out of saliva. The Texas cave salamander has neither eyes nor pigment. Unique access to a hidden world of stalactites, stalagmites, snotites and troglodytes brings a wealth of surprises.

Cities are growing at a faster rate than any other habitat on Earth. They may seem an unlikely place for animals to thrive, but they can be a world of surprising opportunity. Leopards prowl the streets of Mumbai, peregrine falcons hunt amongst New York's skyscrapers, and a million starlings perform spectacular aerial dances over Rome. In Jodhpur, langurs are revered as religious deities and in Harar, locals live in harmony with wild hyenas. Many animals, however, struggle to cope in the urban jungle. As the architects of this environment, can humans choose to build cities that are homes for both them and wildlife?

Japan's landscapes range from snowy mountains to subtropical warmth. They are full of wildlife - and animals' and people's lives often cross as they adapt to these extremes. The central island of Honshu is home to over 100 million people, and its biggest city, Tokyo, is one of the largest urban metropolises on earth. But it has a wild heart - most of Honshu is mountainous. This wilderness is home to an astonishing range of wildlife - black bears, monkeys, exquisite fireflies and even cow demons. But all across this island, from the mountains to the edge of the sea, people and nature are drawn together in the most unexpected ways.

From flightless cormorants hunting underwater to giant tortoises courting on the rim of an active volcano, a look at the hidden side of Galapagos, revealing why it is such a fascinating showcase for evolution.

David Attenborough concludes his epic history of the evolution of flight with an exploration of the highly advanced fliers that dominate our skies today – the extraordinarily diverse skills of the birds, and the sonar-guided precision of the bats. He encounters some of our planet’s most remarkable fliers – from peregrine falcons dive-bombing starlings over Rome, to hummingbirds hovering in the cloud forests of Ecuador. In a spectacular finale, he visits Gomantong cave in Borneo to witness the mass exodus of a million bats.