A visit to arguably the most famous archipelago on Earth, the Galapagos. It's home to a myriad of bizarre and unique creatures, endemic to these islands - but how did they get here and what is the key to these extraordinary islands that allows them to thrive? The programme reveals that this key holds not just the secret to life here, but also to how Darwin was able to leave with the ideas that would revolutionise biology.

The Cave of Swallows in Mexico is a 400m vertical shaft, deep enough to engulf the Empire State Building. The Lechuguilla cave system in the USA is 193km long and 500m deep with astonishing crystal formations hanging from its chambers. Although often overlooked, caves are remarkable habitats with equally bizarre wildlife. Cave angel fish cling to the walls behind cave waterfalls with microscopic hooks on their flattened fins. Cave swiftlets navigate by echo-location and build nests out of saliva. The Texas cave salamander has neither eyes nor pigment. Unique access to a hidden world of stalactites, stalagmites, snotites and troglodytes brings a wealth of surprises.

Around 30% of the land's surface is desert, the most varied of our ecosystems despite the lack of rain. Unravel the secrets of desert survival and experience the ephemeral nature of this dynamic environment. Watch Saharan sandstorms nearly a mile high and desert rivers that run for a single day. In the Gobi Desert, rare Bactrian camels get moisture from the snow. In the Atacama, guanacos survive by licking dew off cactus spines. In the USA, the brief blooming of Death Valley triggers a plague of locusts 65km wide and 160km long. A unique aerial voyage over the Namibian desert reveals elephants on a long trek for food and desert lions searching for wandering oryx.

East Africa is a land which is constantly changing. To survive here, creatures must be able to deal with unpredictable twists and turns - wet turning to dry, feast to famine, cold to hot - no matter how hostile it becomes. From dense forests to snow capped peaks, steamy swamps and endless savannah, this unique and varied land is also a haven for life, supporting large animals in numbers found nowhere else on Earth. But away from the familiar, forever-travelling herds, there are a huge cast of other characters - lizards that steal flies from the faces of lions, vast dinosaur-like birds who stalk catfish through huge wetlands, and an eagle who risks everything on the arrival of ten million bats from a far off rainforest.

The Russian Arctic is one of the world’s most extreme habitats, yet it is a haven for polar bears, lemmings, arctic foxes and walruses. In late August, massive muskoxen bulls gather together to compete for mating dominance. Each bull can weigh 400 kilogram’s and charge at a speed of 40 kilometres an hour; a head-to-head impact can be heard more than a kilometre away

From flightless cormorants hunting underwater to giant tortoises courting on the rim of an active volcano, a look at the hidden side of Galapagos, revealing why it is such a fascinating showcase for evolution.