The contests between predators and prey are the most dramatic events in nature. Taking an intimate and detailed look at the remarkable strategies employed by hunters to catch their prey and the hunted to escape. Sir David Attenborough narrates. "The Hardest Challenge" reveals the extraordinary range of techniques predators use to catch their prey - from a leopard using all its powers of stealth to stalk impala in broad daylight to wild dogs, whose tactic is to wear down their prey over long distances; from Nile crocodiles, the planet's most patient predators, to killer whales who use teamwork and intelligence to take on humpback whales. But even with these finely tuned strategies, the outcome is far from certain. Surprisingly, most predators fail most of the time.

A humpback whale mother and calf embark on an epic journey from tropical coral paradises to storm ravaged polar seas. Newly discovered coral reefs in Indonesia reveal head-butting pygmy seahorses, flashing 'electric' clams and bands of sea kraits, 30-strong, which hunt in packs. Elsewhere plagues of sea urchins fell forests of giant kelp. Huge bull fur seals attack king penguins, who despite their weight disadvantage, put up a spirited defence.

Mammals dominate the planet. They do it through having warm blood and by the care they lavish on their young. Weeks of filming in the bitter Antarctic winter reveal how a mother Weddell seal wears her teeth down keeping open a hole in the ice so she can catch fish for her pup. A powered hot air balloon produces stunning images of millions of migrating bats as they converge on fruiting trees in Zambia, and slow-motion cameras reveal how a mother rufous sengi exhausts a chasing lizard. A gyroscopically stabilised camera moves alongside migrating caribou, and a diving team swim among the planet's biggest fight as male humpback whales battle for a female.

Our journey begins with David Attenborough at the North Pole, as the sun returns after six months of darkness. We follow a pair of courting polar bears, which reveal a surprisingly tender side. Next stop is the giant Greenland ice cap, where waterfalls plunge into the heart of the ice and a colossal iceberg carves into the sea. Humpback whales join the largest gathering of seabirds on earth to feast in rich Alaskan waters. Further south, the tree line marks the start of the Taiga forest, containing one third of all trees on earth. Here, 25 of the world's largest wolves take on formidable bison prey. At the other end of our planet, the Antarctic begins in the Southern Ocean where surfing penguins struggle to escape a hungry sea-lion and teams of orcas create giant waves to wash seals from ice floes -a filming first. Diving below the ice, we discover prehistoric giants, including terrifying sea spiders and woodlice the size of dinner plates. Above ground, crystal caverns ring the summit of Erebus, the most southerly volcano on earth. From here we retrace the routes of early explorers across the formidable Antarctic ice-cap - the largest expanse of ice on our planet. Finally, we rejoin David at the South Pole, exactly one hundred years after Amundsen then Scott were the first humans to stand there

Few sounds are more beautiful or moving than the underwater songs of the humpback whale. Male whales compete with their songs, which often last for 10 minutes at a time, and can be repeated for hours on end. Whales separated by thousands of miles of sea will sing almost identical songs. Researchers have found that the songs change throughout the breeding months, following a mysterious pattern repeated across the waves. Whales also use sound to hunt. To catch herring, humpback whales release a stream of bubbles to form a shimmering, circular fishing net. Emitting a repetitive loud scream, they scare the fish into a tight ball, then lunge out of the water to swallow the shoal whole. Now it seems that the long-held image of the gentle giant must change to one of a ferocious and opportunistic hunter.