One hundred years after Columbus' arrival in the Caribbean, Spanish Conquistadors and Priests, push into North America in search of gold and to spread Catholicism. With the arrival of the British in North America, the two colonial systems produce contrasting societies that come in conflict as Manifest Destiny pushes the U.S into the Mexican territories of the South West. As the Gold Rush floods California with settlers, complex and vital communities are overwhelmed. The elites, including Mariano Vallejo and Apolinaria Lorenzana lose their land. Mexicans and Mexican Americans are treated as second-class citizens, facing discrimination and racial violence. Resistance to this injustice appears in New Mexico as Las Gorras Blancas (The White Caps), burn Anglo ranches and cut through barbed wire to prevent Anglo encroachment. At the same time, New Mexicans manage to transform themselves through education, managing to preserve Hispano culture in New Mexico and their standing in the midst of an era of conquest and dispossession.

World War II is a watershed event for Latino Americans with hundreds of thousands of men and women serving in the armed forces, most fighting side by side with Anglos. In the Pacific, East L.A.'s Guy Gabaldon becomes a Marine Corp legend when he singlehandedly captures more enemy soldiers than anyone in US military history. But on the home front, discrimination is not dead: in 1943, Anglo servicemen battle hip young "Zoot suitors" in racially charged riots in southern California

In the 1960s and 1970s a generation of Mexican Americans, frustrated by persistent discrimination and poverty, find a new way forward, through social action and the building of a new "Chicano" identity. The movement is ignited when farm workers in the fields of California, led by Cesar Chavez and Dolores Huerta, march on Sacramento for equal pay and humane working conditions. Through plays, poetry and film, Luis Valdez and activist Corky Gonzalez create a new appreciation of the long history of Mexicans in the South West and the Mestizo roots of Mexican Americans. In Los Angeles, Sal Castro, a schoolteacher, leads the largest high school student walkout in American history, demanding that Chicano students be given the same educational opportunities as Anglos. In Texas, activists such as Jose Angel Gutierrez, create a new political party and change the rules of the electoral game. By the end of the 1970s Chicanos activism and identity have transformed what it means to be an American. Chicano and Latino studies are incorporated into school curriculum; Latinos are included in the political process.

Dr Janina Ramirez unlocks the secrets of illuminated manuscripts that were custom-made for kings and explores the medieval world they reveal. 'Ruling by the Book.' Janina begins her journey with the first Anglo-Saxon rulers to create a united England, encountering books in the British Library's Royal manuscripts collection which are over a thousand years old and a royal family tree which is five metres long.
Janina finds out about a king who had a reputation for chasing nuns and reads a book created as a wedding gift for a ten-year-old prince. She roams from Westminster Abbey to other ancient English spiritual sites such as Winchester, St Albans and Malmesbury, and sees for herself how animal skins can be transformed into the finest vellum.

In a late Jurassic forest in what is now China, an Epidexipteryx escapes from a juvenile Sinraptor by climbing a tree. It finds a beetle grub in the tree bark, being shown to use its elongated fingers in a similar way to a modern day aye-aye. However, its prey is stolen by another, larger Epidexipteryx, and after a brief bout of posturing, the smaller individual goes to find more food. It drops a second grub to the forest floor, where the other Epidexipteryx retrieves it, only to be killed by the juvenile Sinraptor. The episode then cuts to a desert in late Cretaceous Mongolia, where a Saurornithoides is shown brooding a nest of eggs. When it leaves the nest, an Oviraptor raids it, fleeing when the troodontid returns. The Saurornithoides is suddenly attacked and eaten by a Gigantoraptor, which then heads to compete in a breeding ritual for mates. The males use their feathers for display, a brief fight between two erupting at one point, allowing the females to choose the best suitor. The episode finally cuts to an early Cretaceous forest in China, where a Xianglong is being hunted by a Microraptor, which uses its feathers to pursue the gliding lizard in the air. A Sinornithosaurus attacks it, and after a brief chase the Microraptor manages a lucky escape. The Sinornithosaurus, alongside two other members of its species is then shown hunting a Jeholosaurus and its three young. The group brings down the parent, the narrator explaining that their possibly venomous bite allowed them to tackle animals much larger than themselves. A montage is then shown of the feathered dinosaurs featured in the programme, with the narrator saying that Microraptor not only hints at how flight might have developed, but also that dinosaurs still live amongst us today, as birds.

Professor Robert Bartlett explores how the Normans developed from a band of marauding Vikings into the formidable warriors who conquered England in 1066. He tells how the Normans established their new province of Normandy -'land of the northmen' - in northern France. Under the leadership of Duke William, they conquered England. The Battle of Hastings marked the end of the Anglo-Saxon aristocracy and monarchy. The culture and politics of England would now be transformed by the Normans.