Africa's wildest river is home to the most spectacular wildlife. Hippos fight for territory while herds of elephant, water buffalo and zebra depend on it for life. In the wet season the rains burst the riverbanks and everyone, including people, must move whilst fish swim through the villages. In the dry season the creatures fight over the few pools of water while predators prowl. At its heart it plunges over Victoria Falls and into wild ravines before draining into the Indian Ocean, where storm clouds cycle the water back into the heart of Africa.

Reptiles and amphibians look like hang-overs from the past. But they overcome their shortcomings through amazing innovation. The pebble toad turns into a rubber ball to roll and bounce from its enemies. Extreme slow-motion shows how a Jesus Christ lizard runs on water, and how a chameleon fires an extendible tongue at its prey with unfailing accuracy. The camera dives with a Niuean sea snake, which must breed on land but avoids predators by swimming to an air bubble at the end of an underwater tunnel. In a TV first, Komodo dragons hunt a huge water-buffalo, biting it to inject venom, then waiting for weeks until it dies. Ten dragons strip the carcass to the bone in four hours.

The great flood in the Okavango turns 4,000 square miles of arid plains into a beautiful wetland. Elephant mothers guide their families on an epic trek across the harsh Kalahari Desert towards it, siphoning fresh water from stagnant pools and facing hungry lions. Hippos battle for territory, as the magical water draws in thousands of buffalo and birds, and vast clouds of dragonflies. Will the young elephant calves survive to reach this grassland paradise? The experienced mother elephants time their arrival at the delta to coincide with the lush grass produced by the great flood.
In a TV first, the programme shows the way they use their trunks to siphon clean water from the surface layers of a stagnant pool, while avoiding stirring up the muddy sediment on the bottom with their feet. Lechwe swamp deer, zebras, giraffes, crocodiles and numerous fish and thousands of birds arrive in the delta. And, in a phenomenon never before filmed in the Okavango, thousands of dragonflies appear - seemingly from nowhere - within minutes of the flood arrival, mating and laying eggs. As the flood finally reaches its peak, elephants and buffalo, near the end of their epic trek across the desert, face the final gauntlet of a hungry pride of lions. In a heart-wrenching sequence, a baby elephant is brought down by a lion in broad daylight.