As the early June sun begins to set over a calm river in Central Hungary, masses of ghostly shapes emerge from their larval cases to take to the air for the first time. They are mayflies and in a spectacular display, thousands of them demonstrate how the very first wings were used. From the stunning aerobatics of hoverflies in an English garden to the mass migration of purple crow butterflies in the valleys of Taiwan, this episode tells the tale of the first animals ever to take to the air. Unique footage reveals the lightning fast reactions of bluebottles and hoverflies, filmed with one of the world's fastest cameras, and Sir David Attenborough handles the world's largest (and perhaps most ferocious) insect - the Titan beetle.

Silk is the invertebrates' great invention, used in a range of ways from from the protective stalks of lacewing eggs to the amazing hanging threads of New Zealand's 'glow worms'. Spiders, though, have taken silk-spinning to extremes. The common wolf spider has no web, but the female is a gentle parent, encasing her eggs in silk and carrying the precious bundle wherever she goes. The bolas spider uses a ball of sticky silk soaked in a copy of moth pheromone to lure its prey. Millions of communal spiders live and feed together in a vast, towering web - an arachnophobe's nightmare.

The world of invertebrates exists in a web of relationships with plants and other animals. Unique footage of the world's smallest insect (a fairy wasp only quarter of a millimetre long) shows it flying underwater to find the eggs of water beetles in which to lay its own brood. Some ants 'farm' the trees that give them shelter, creating areas known as 'Devil's gardens'. To make sure these grow without competition, they kill off other seedlings in the surrounding vegetation. The blister beetle's larvae huddle together on the end of a piece of grass and mimic a female bee. When a male bee tries to mate with the 'female', the larvae grab on to his belly. Confused, he flies away and searches for a real female. When he eventually finds her and mates with her, the beetle larvae hurriedly swap from his front on to her back, and hence get carried back to her nest where they eat her pollen supplies.

Invertebrates don't always operate alone. True society was the last feature to evolve in invertebrates, as recently as the time of Tyrannosaurus. In the last programme see the tensions below the surface in some of the great social structures built by insects, and witness the carnage when an ant colony and a termite colony wage war.

After eight grueling years of hunting in the hot, wind-scoured desert of central Africa, an international team of researchers has uncovered one of the most sensational fossil finds in living memory: the well-preserved 7 million years old skull of a chimp-size animal, probably a male, that doesn't fit any known species. According to paleontologist Michel Brunet of the University of Poitiers in France, whose team reported the find in Nature last week, there is no way it could have been an ape of any kind. It was almost certainly a hominid — a member of a subdivision of the primate family whose only living representative is modern man.

It was the bloodiest war in history, yet Hitler's brutal invasion of the Soviet Union and Stalin's subsequent fight back is surprisingly little known in the West. More than 30 million people died during Hitler and Stalin's terrible war in the East - more than in any other single war in history. With direct testimony from people who perpetrated atrocities and the victims who suffered on both sides, we bring the unspeakable to the screen and allowing victims and aggressors to tell the truth as they see it. In the first episode, we will look at Hitler's plans to invade the Soviet Union and the horror of the first months of conflict.