Nature

A dramatic journey along the coasts of South America, from the Falklands to the tropical Galapagos. The cold currents create rich fisheries, and support such diverse marine life as dolphins, killer whales, sea lions and albatross. But the most biquitous creatures are the penguins, which inhabit surprising locations - lush forest, arid desert and even equatorial areas.

They soar through the heavens, fly through the oceans and glide along land. But these are not creatures found on a wildlife safari. These are life forms from another planet. Armed with scientific fact and a little imagination, experts come together to take you on an unprecedented journey to the edges of our imagination.

As the early June sun begins to set over a calm river in Central Hungary, masses of ghostly shapes emerge from their larval cases to take to the air for the first time. They are mayflies and in a spectacular display, thousands of them demonstrate how the very first wings were used. From the stunning aerobatics of hoverflies in an English garden to the mass migration of purple crow butterflies in the valleys of Taiwan, this episode tells the tale of the first animals ever to take to the air. Unique footage reveals the lightning fast reactions of bluebottles and hoverflies, filmed with one of the world's fastest cameras, and Sir David Attenborough handles the world's largest (and perhaps most ferocious) insect - the Titan beetle.

The world of invertebrates exists in a web of relationships with plants and other animals. Unique footage of the world's smallest insect (a fairy wasp only quarter of a millimetre long) shows it flying underwater to find the eggs of water beetles in which to lay its own brood. Some ants 'farm' the trees that give them shelter, creating areas known as 'Devil's gardens'. To make sure these grow without competition, they kill off other seedlings in the surrounding vegetation. The blister beetle's larvae huddle together on the end of a piece of grass and mimic a female bee. When a male bee tries to mate with the 'female', the larvae grab on to his belly. Confused, he flies away and searches for a real female. When he eventually finds her and mates with her, the beetle larvae hurriedly swap from his front on to her back, and hence get carried back to her nest where they eat her pollen supplies.

Invertebrates don't always operate alone. True society was the last feature to evolve in invertebrates, as recently as the time of Tyrannosaurus. In the last programme see the tensions below the surface in some of the great social structures built by insects, and witness the carnage when an ant colony and a termite colony wage war.

The Russian Arctic is one of the world’s most extreme habitats, yet it is a haven for polar bears, lemmings, arctic foxes and walruses. In late August, massive muskoxen bulls gather together to compete for mating dominance. Each bull can weigh 400 kilogram’s and charge at a speed of 40 kilometres an hour; a head-to-head impact can be heard more than a kilometre away