The contests between predators and prey are the most dramatic events in nature. Taking an intimate and detailed look at the remarkable strategies employed by hunters to catch their prey and the hunted to escape. Sir David Attenborough narrates. "The Hardest Challenge" reveals the extraordinary range of techniques predators use to catch their prey - from a leopard using all its powers of stealth to stalk impala in broad daylight to wild dogs, whose tactic is to wear down their prey over long distances; from Nile crocodiles, the planet's most patient predators, to killer whales who use teamwork and intelligence to take on humpback whales. But even with these finely tuned strategies, the outcome is far from certain. Surprisingly, most predators fail most of the time.

Silk is the invertebrates' great invention, used in a range of ways from from the protective stalks of lacewing eggs to the amazing hanging threads of New Zealand's 'glow worms'. Spiders, though, have taken silk-spinning to extremes. The common wolf spider has no web, but the female is a gentle parent, encasing her eggs in silk and carrying the precious bundle wherever she goes. The bolas spider uses a ball of sticky silk soaked in a copy of moth pheromone to lure its prey. Millions of communal spiders live and feed together in a vast, towering web - an arachnophobe's nightmare.

the final episode deals with plants that live in hostile environments. Attenborough visits Ellesmere Island, north of the Arctic Circle, to demonstrate that even in a place that is unconducive to life, it can be found. Algae and lichens grow in or on rock, and during summer, when the ice melts, flowers are much more apparent. However, they must remain close to the ground to stay out of the chilling wind. In the Tasmanian mountains, plants conserve heat by growing into 'cushions' that act as solar panels, with as many as a million individual shoots grouped together as one. Others, such as the lobelia in Mount Kenya, have a 'fur coat' of dense hairs on their leaves. The saguaro cactus in the Sonoran Desert flourishes because of its ability to retain vast amounts of water, which can't be lost through leaves because it has none. Many desert dwellers benefit from an accelerated life cycle, blooming rapidly within weeks after rainfall. Conversely, Mount Roraima is one of the wettest places on Earth. It is a huge sandstone plateau with high waterfalls and nutrients are continuously washed away, so plants have to adapt their diet if they are to survive. A bladderwort is shown invading a bromeliad. Inhabitants of lakes have other problems to contend with: those that dominate the surface will proliferate, and the Amazon water lily provides an apt illustration. Attenborough ends the series with an entreaty for the conservation of plant species.

Reptiles and amphibians look like hang-overs from the past. But they overcome their shortcomings through amazing innovation. The pebble toad turns into a rubber ball to roll and bounce from its enemies. Extreme slow-motion shows how a Jesus Christ lizard runs on water, and how a chameleon fires an extendible tongue at its prey with unfailing accuracy. The camera dives with a Niuean sea snake, which must breed on land but avoids predators by swimming to an air bubble at the end of an underwater tunnel. In a TV first, Komodo dragons hunt a huge water-buffalo, biting it to inject venom, then waiting for weeks until it dies. Ten dragons strip the carcass to the bone in four hours.

Marine invertebrates are some of the most bizarre and beautiful animals on the planet, and thrive in the toughest parts of the oceans. Divers swim into a shoal of predatory Humboldt squid as they emerge from the ocean depths to hunt in packs. When cuttlefish gather to mate, their bodies flash in stroboscopic colours. Time-lapse photography reveals thousands of starfish gathering under the Arctic ice to devour a seal carcass. A giant octopus commits suicide for her young. A camera follows her into a cave which she walls up, then she protects her eggs until she starves. The greatest living structures on earth, coral reefs, are created by tiny animals in some of the world's most inhospitable waters.

Fifty years ago there were close to half-a-million lions in Africa. Today there are around 20,000. To make matters worse, lions, unlike elephants, which are far more numerous, have virtually no protection under government mandate or through international accords. This is the jumping-off point for a disturbing, well-researched and beautifully made cri de coeur from husband and wife team Dereck and Beverly Joubert, award-winning filmmakers from Botswana who have been Explorers-in-Residence at National Geographic for more than four years. Pointing to poaching as a primary threat while noting the lion's pride of place on the list for eco-tourists-an industry that brings in 200 billion dollars per year worldwide-the Jouberts build a solid case for both the moral duty we have to protect lions (as well as other threatened "big cats," tigers among them) and the economic sense such protection would make. And when one takes into account the fact that big cats are at the very top of the food chain-and that their elimination would wreak havoc on all species below them, causing a complete ecosystem collapse-the need takes on a supreme urgency.