A dramatic journey along the coasts of South America, from the Falklands to the tropical Galapagos. The cold currents create rich fisheries, and support such diverse marine life as dolphins, killer whales, sea lions and albatross. But the most biquitous creatures are the penguins, which inhabit surprising locations - lush forest, arid desert and even equatorial areas.

From flightless cormorants hunting underwater to giant tortoises courting on the rim of an active volcano, a look at the hidden side of Galapagos, revealing why it is such a fascinating showcase for evolution.

Animals and plants in the Galapagos have evolved the most surprising ways to cope with the profound geological and climatic forces.

The next instalment focuses on dietary needs and how different species have evolved beaks to suit their individual requirements. The latter come in a multitude of forms. Blue tits and goldfinches have beaks akin to tweezers, with which to extract seeds, while the hawfinch's razor-like bill can deal with a cherry-stone. However, the crossbill is the only finch that can twist its mandibles in opposite directions. Jays store acorns for winter by burying them in the ground, whereas woodpeckers can keep up to 60,000 of them in one tree trunk. Sap is also desirable, and there are a variety of methods used to obtain it. The hoatzin is the only specialised leaf-eater, and accordingly has a digestive system more akin to that of cattle. Plants recruit birds to aid pollination, and offer nectar as a reward. Hummingbirds eat little else, and the sword-bill's beak is the longest of any bird in relation to its body. Insects are also highly prized, and Galapagos finches are shown to possess some ingenuity as they not only strip bark, but also use 'tools' to reach their prey. Crows are hailed as being among the most intelligent birds, and one is shown using a twig to spear a grub within a fallen log. The robin is an opportunist, and Sir Attenborough observes one seizing morsels as he digs a patch of earth. In South America, a cattle tyrant sits atop an obliging capybara and uses its vantage point to spot passing food that may be dislodged by its grazing partner.

Once life arrived in the Galapagos, it exploded into unique and spectacular forms. David Attenborough investigates the driving forces behind such evolutionary innovations. We learn that life must be able to adapt quickly in these ever-changing volcanic landscapes. It has resulted in species found nowhere else in the world, such as giant whale sharks and marine iguanas that can spit sea-salt from their noses, dandelion seeds that grow into tree-sized plants and spiders that can blend perfectly into the darkness. Adaptation has been the key to survival in these islands so far, but the story of life in the Galapagos doesn’t end here. The catalyst that triggers these explosions of life remains in place.

No two islands in the Galapagos are the same. The imperceptible drift of a continental plate keeps each island biologically isolated. David Attenborough explores this evolutionary crucible, encountering tortoises that weigh up to half a tonne, finches that use tools and lizards that communicate using press-ups; for Darwin, this was all evidence for his theory of evolution. We see the final footage of the world famous tortoise fondly known as Lonesome George, the last survivor of his species. David Attenborough was the last person to have ever filmed with him. Darwin’s famous visit had a downside – the arrival of man. David investigates the impact we’ve had in these islands, as our influence is a double-edged sword. We’ve disrupted the natural balance but he also believes Darwin would be thrilled with the advances we have made in science. We’re also now uncovering evidence that evolution is more rapid than Darwin could ever have imagined. Whatever wonders the Galapagos Islands hold today, they are only a hint of what awaits them in the future.