On the coast, two worlds collide. Coasts are the most dynamic and challenging habitats in the ocean - that brings great rewards but also great danger. The extraordinary animals that live here must find ingenious ways to cope with two very different worlds. This episode is a rollercoaster ride of heart-stopping action and epic drama, peopled with characters from the beautiful to the bizarre. We meet fish that live on dry land and puffins that must travel 60 miles or more for a single meal, and witness a life-and-death struggle in a technicolour rock pool. In a secluded cove in the Galapagos, sea lions feast on 60kg tuna. It should be impossible - tuna are usually far too fast for sea lions to catch. But here the sea lions club together to herd their prey inshore. Once trapped in the shallows, these huge fish are easy pickings. As the tide recedes in Brazil, lightfoot crabs leap from rock to rock, desperately avoiding the water - their lives depend on it. Moray eels launch themselves from rock pools, jaw gaping. Then octopuses, too. Both crawl across dry rock to set their ambush. Elsewhere, the ever-changing tides create rock pools. But these temporary worlds are a battleground. Predatory starfish turn a magical garden into the stuff of nightmares.

The years 1825-1918 were bloody and traumatic, a period when four tsars tried - and failed - to deal with the growing pressure for constitutional reform and revolution. In 1861, millions of enslaved serfs were freed by the Tsar-Liberator, Alexander II. But Alexander paid the ultimate penalty for opening the Pandora's box of reform when he was later blown up by terrorists on the streets of St Petersburg. Elsewhere, there was repression, denial, war and - in the case of the last tsar, Nicholas II - a fatalistic belief in the power of God, with Nicholas's faith in the notorious holy man Rasputin being a major part in his undoing. Lucy Worsley also details the chilling murder of Nicholas and his family in 1918, and asks whether all of this horror have been avoided.

Episode three explores the mind of one of the most fanatical of all Nazis and the insight that gives into the psychology of dictatorship. Hess is brought to Nuremberg from the UK where he had flown 4 years earlier much to the bemusement of the British. Overy explains that they must have quickly realised he was not normal. In the intervening time Hess’s mental state has further deteriorated and when interviewed by Chief Interrogator Colonel John Amen proclaims amnesia. Chief Interpreter Richard Sonnenfeldt explains that they brought in Göring to confront Hess but that he too failed to make an impression.
Psychopathologist Prof. Edgar Jones proposes Hess’s behaviour patterns may be his way of escaping reality as he is forced to face the extent of the atrocities and choose between accepting his part of the blame or forsaking his Führer. However, as history Professor Robert Gellatel explains it is difficult to construct a case against Hess as he was imprisoned in England when the worst of the atrocities were carried out so British Prosecutor Mervyn Griffith-Jones must argue a conspiracy charge linking the pre-1939 persecutions to the post-1939 atrocities and a crimes against peace charge proving the flight to Scotland was merely a ruse.

David Attenborough reveals the fascinating lives of snakes, the most misunderstood group of reptiles. A CCTV stakeout of wild rattlesnakes hunting shows, for the first time, what sophisticated predators they truly are. Attenborough is also on the receiving end of a spitting cobra's chemical weapons system, while the surprisingly beautiful and tender side of snakes is displayed in the sinuous courtship of kingsnakes, and the water birth of 15 beautiful yellow anaconda babies.

Dr Janina Ramirez shows how medieval manuscripts gave power to the king and united the kingdom in an age of plague, warfare and rebellion, discovers that Edward III used the manuscripts he read as a boy to prepare him for his great victory at the battle of Crecy and reveals how a vigorous new national identity bloomed during the 100 Years War with France. In the British Library's Royal Manuscripts collection Dr Ramirez finds out that magnificent manuscripts like the Bedford Hours, taken as war booty from the French royal family, were adapted for the education of English princes. She also explores how knowledge spread through a new form of book - the encyclopaedia.

The story of the British Library's Royal Manuscripts collection reaches its end with the last great flowering of illumination, in the magnificent courts of the Tudors. Dr Janina Ramirez investigates astrological texts created for Henry VII, and unwraps his will - still in its original, extravagantly-decorated velvet and gold cover. She hears music written for Henry VIII, which went unperformed for centuries; and reads love notes between the king and Anne Boleyn, written in the margins of a prayer book. Nina also visits Bruges, the source of many of the greatest manuscripts, where this medieval art form collided with the artistic innovations of the Renaissance.