The Arctic is the harshest environment on Earth: little food grows, it's dark for months on end, and temperatures stay well below freezing for much of the year. Yet four million people manage to survive here. Human Planet tells remarkable stories of extraordinary people who make their homes in nature's deep freeze. In springtime, Amos and Karl-Frederik set out across the sea ice with their dogs to catch a real-life sea monster: a Greenland shark! Inuit mussel-gatherers venture underneath the sea ice at low tide for a perilous race against time as they gather their food. And the children of Churchill, Manitoba, set out on the most dangerous trick or treating Halloween in the world: they risk coming face-to-face with deadly polar bears on the streets of their town. Who'll get the tastiest snack?

This episode discusses the human exploration of Antarctica, in particular the mission led by Captain Robert Falcon Scott, whose team died on the way back from the South Pole. It shows the scientific work in the modern human bases in Antarctica, especially Mawson Base and its observation of Adelie Penguins (partially through tracking devices). The second half of the episode describes how the series was made. Most crucial was of course the camera work. To get access to the wildlife of the sea, for example, boats, divers, suspended capsules and remotely controlled cameras mounted on inflatables were used. Particularly dangerous to divers were Leopard Seals and other predators. The film concludes that although working in Antarctica is now much easier than during the early days of exploration, human footsteps on the continent are still exceedingly rare in part because of international treaties prohibiting industrial exploitation.

Are the wealthy just born in the right place at the right time? Are the poor victims of a system designed to keep them down? Or do physics and biology determine who is rich and who is poor? Throughout history, distribution of wealth is governed by hidden forces: DNA, environmental stress, patterns of human migration and even the laws of thermodynamics! Nature seems to demand winners and losers in life. But does this mean greed is king, and the rich can take what they want? Or is cooperation – between microorganisms, monkeys, and humans – more essential to survival of a species?

In the final challenge of the game of life, raising offspring is the ultimate prize. Continuing the line through the next generation, is the next best thing to immortality. But it’s far from easy... Some parents must risk their own lives for their offspring.

Brian travels across the US and encounters some astonishing creatures that reveal how the senses evolved. It is a story that takes him through life's journey, from single-celled organisms to sentient beings.

In this final episode we complete our journey, travelling back from the March equinox to the end of June. Kate Humble is in the Arctic at a place where spring arrives with a bang, whilst Helen Czerski chases a tornado to show how the earth's angle of tilt creates the most extreme weather on earth.