The big blue is the world's greatest wilderness, far from shore and many kilometres deep. It's a vast marine desert where there is little to eat and nowhere to hide. Yet it's home to some of the biggest and most spectacular creatures on earth. This episode reveals what it takes to survive in this savage and forbidding world. We witness feats of incredible endurance, moments of high drama and extraordinary acts of heart-wrenching self-sacrifice. Every animal in the big blue must find their own unique way to survive. Sperm whales have the largest brains in the world. They live for 80 years, and we are only now beginning to learn the extraordinary complexity of their language of clicks - thought to coordinate the whole family in everything from childcare to hunting. With special pressure-proof cameras, we witness record-breaking feats of endurance as they hunt for squid a kilometre down into the abyss. Only recently have we begun to solve the mystery of where baby turtles disappear to in their early years. They leave the crowded waters of the coast and head to the open ocean, where they use floating debris like logs as life rafts. Here they remain until adulthood, adrift on the high seas in relative safety away from coastal predators. Over half of all animals in the open ocean drift in currents. Jellyfish cross entire oceans feeding on whatever happens to tangle with their tentacles. The jelly-like Portuguese man-of-war can harness sail power to fish with its deadly tentacles. Sometimes there is a brief explosion of food in this marine desert, but ocean hunters must be fast to make the best of this bonanza. We witness super pods of up to 5,000 spinner dolphins racing to herd vast shoals of lanternfish, briefly caught at the surface where it is thought they spawn. New aerial footage reveals, for the first time, the truth to a centuries-old sailors' legend of the 'boiling seas' - the spectacular feeding frenzy of 90kg tuna and dolphins smashing through the lantern fish shoals turning the sea white with foam.

In the second episode, the directors learn their vision for the films is not always permissible by the U.S. government. Wyler is shocked by the racism he encounters against African American soldiers and refused to make a film recruiting black soldiers. Meanwhile, the films' racist depiction of the Japanese versus human depiction of the Germans causes worry for the War Department, which at that time planned to redistribute the Japanese-American population from internment camps into towns across the United States.

The five directors, John Ford, William Wyler, John Huston, Frank Capra, and George Stevens, return to Hollywood after the war but are forever haunted by what they saw. Ford goes on a drinking bender after filming the carnage at D-Day. Stevens is wholly unprepared for the horrors of Dachau and realizes he is not there to film propaganda but to capture evidence of crimes against humanity. Wyler, who lost his hearing during the war, fears his career is over. Huston chronicles soldiers suffering from postraumatic stress disorder in the film Let There Be Light, only to have it suppressed by the U.S. government.

Award winning wildlife photographers Jonathan and Angela Scott have travelled the world, always returning home to Kenya. There they introduced the world to the big cats of the Maasai Mara and now they are part of its fight for survival.

Africa's wildest river is home to the most spectacular wildlife. Hippos fight for territory while herds of elephant, water buffalo and zebra depend on it for life. In the wet season the rains burst the riverbanks and everyone, including people, must move whilst fish swim through the villages. In the dry season the creatures fight over the few pools of water while predators prowl. At its heart it plunges over Victoria Falls and into wild ravines before draining into the Indian Ocean, where storm clouds cycle the water back into the heart of Africa.

Lucy Worsley continues her journey through Russia in the footsteps of the Romanovs, the most powerful royal dynasty in modern European history. In this episode she examines the extraordinary reign of Catherine the Great, and the traumatic conflict with napoleonic France that provides the setting for the novel War and Peace. At the magnificent palace of Peterhof near St Petersburg, Lucy charts the meteoric rise of Catherine the Great, who seized the Russian throne from her husband Peter III in 1762 and became the most powerful woman in the world. Catherine was a woman of huge passions - for art, for her adopted country (she was German by birth) and for her many lovers.
Catherine expanded her empire through military victories overseas, while at home she encouraged education and introduced smallpox inoculation to Russia. But Catherine struggled to introduce deeper reforms, and the institution of serfdom remained largely unchanged. Lucy explains how this injustice fuelled a violent rebellion. Nevertheless, Catherine left Russia more powerful on the world stage than ever. But all she had achieved looked set to be undone when Napoleon invaded Russia in 1812. Lucy relives the pivotal battle of Borodino, when the Russian army finally confronted the French forces; the traumatic destruction of Moscow; and, under Catherine's grandson Alexander, the eventual victory over the French that provided the Romanov dynasty with its most glorious hour.