Bladderwort utricularia is a pond-dweller that is among the fastest known, its traps snapping shut in less than a millisecond. As the seasons change, David demonstrates how plants operate on a different time scale to us; how they modify their lives according to the time of year. We discover insects’ hidden links with plants, both as pests and pollinators. UV-sensitive 3D cameras reveal the invisible alter-ego of plants and their flowers’ mesmerizing patterns; a parallel-dimension of strange colours and stunning patterns through which plants communicate with them. With the aid of visual effects, David steps among the swirling vortices of plant scent; communication signals with which plants are inextricably plugged in to the natural world. And using a tuning fork, he demonstrates how plants and insects can even communicate with music. As autumn envelopes the Gardens, fungi reveal themselves not as the enemies of plants but their vital allies. In Kew’s atmospheric Fungarium, David discovers a specimen that has the power of mind control and another that lives underground where it has grown to be so big it can be counted as the largest single organism on the planet. It is 6 times bigger than Kew Gardens itself.

This episodes the nature of the cosmos on the micro and atomic scales, using the Ship of the Imagination to explore these realms. Tyson describes some of the micro-organism that live within a dew drop, demonstrating parameciums and tardigrades. He proceeds to discuss how plants use photosynthesis via their chloroplasts to convert sunlight into chemical reactions that convert carbon dioxide and water into oxygen and energy-rich sugars. Tyson then discusses the nature of molecules and atoms and how they relate to the evolution of species. He uses the example set forth by Charles Darwin postulating the existence of the long-tongued Morgan's sphinx moth based on the nature of the comet orchid with pollen far within the flower. He further demonstrates that scents from flowers are used to trigger olfactory centers in the brain, stimulating the mind to threats as to aid in the survival of the species. Tyson narrates how Greek philosophers Thales and Democritus postulated that all matter was made up of combinations of atoms in a large number of configurations, and describes how carbon forms the basic building block for life on earth due to its unique chemical nature. Tyson explains on the basic atomic structure of protons, neutrons, and electrons, and the nature of nuclear fusion that occurs in most stars. He then discusses the existence of neutrinos that are created by these nuclear processes in stars, and that detecting such sub-atomic particles which normally pass through matter require subterranean facilities like the Super-Kamiokande that were used to detect neutrinos from the supernova SN 1987A in the Large Magellanic Cloud before light from the explosion were observed due to their ability to pass through matter of the dying sun. Tyson compares how neutrinos were postulated by Wolfgang Pauli to account for the conservation of energy from nuclear reactions in the same manner as Darwin's postulate on the long-tongued moth. Tyson concludes by noting that there are neutrinos from the Big Bang still existing in the universe but due to the nature of light, there is a "wall of infinity" that cannot be observed beyond.

The fifth programme explores the alliances formed between the animal and plant worlds. Attenborough dives into Australia's Great Barrier Reef and contrasts the nocturnal feeding of coral, on microscopic creatures, with its daytime diet of algae. Some acacias are protected by ants, which will defend their refuge from any predator. Besides accommodation, the guards are rewarded with nectar and, from certain species, protein for their larvae as well. Fungi feed on plants but can also provide essential nutriment to saplings (Mycorrhiza). The connection is never broken throughout a tree's life and a quarter of the sugars and starches produced in its leaves is channelled back to its fungal partners. Meanwhile, fungi that feed on dead wood leave a hollow trunk, which also benefits the tree. Orchids enjoy a similar affiliation. Lichens are the product of a relationship between fungi and a photosynthetic associate, usually algae. They are extremely slow-growing, and a graveyard is the perfect location to discover their exact longevity. Mistletoe is a hemiparasite that obtains its moisture from a host tree, while using own leaves to manufacture food. Its seeds are deposited on another by the mistletoe tyrannulet, following digestion of the fruit. The dodder (Cuscuta) is also parasitic, generally favouring nettles, and siphons its nourishment through periodic 'plugs' along its stem. The rafflesia has no stem or leaves and only emerges from its host in order to bloom — and it produces the largest single flower: one metre across.

Written and presented by David Attenborough, who said: 'One of the most wonderful things about filming plants is that you can reveal hidden aspects of their lives, you can capture the moment as one plant strangles another, and as they burst into flower. But whilst time-lapse photography allows you to see things that no human being has ever seen before". David begins his journey inside the magnificent Palm House, a unique global rainforest in London. Here, he explores the extraordinary plants that are so well adapted to wet and humid environments and unravels the intimate relationships between wet zone plants and the animals that depend on them. It was in the wet zones of the world that plants first moved on to land and in the Waterlily House David reveals how flowers first evolved some 140 million years ago. Watching a kaleidoscope of breath-taking time-lapses of these most primitive of flowers swelling and blooming in 3D, he is able to piece together the very first evolutionary steps that plants took to employ a wealth of insects to carry their precious pollen for the first time. David discovers clues to answer a question that even had Charles Darwin stumped: how did flowering plants evolve so fast to go on to colonise the entire planet so successfully?

In the second episode, Professor Iain Stewart discovers how flowers have transformed our planet. He journeys to the remote islands of the South Pacific to track down the earliest flowers. In the deserts of Africa and rainforests of Vietnam, he sees how they brought brilliant colour to the most barren landscapes and sculpted the earth itself. And he learns how they drove the evolution of all animals - kick-starting our human story.

The story of the British Library's Royal Manuscripts collection reaches its end with the last great flowering of illumination, in the magnificent courts of the Tudors. Dr Janina Ramirez investigates astrological texts created for Henry VII, and unwraps his will - still in its original, extravagantly-decorated velvet and gold cover. She hears music written for Henry VIII, which went unperformed for centuries; and reads love notes between the king and Anne Boleyn, written in the margins of a prayer book. Nina also visits Bruges, the source of many of the greatest manuscripts, where this medieval art form collided with the artistic innovations of the Renaissance.