On the coast, two worlds collide. Coasts are the most dynamic and challenging habitats in the ocean - that brings great rewards but also great danger. The extraordinary animals that live here must find ingenious ways to cope with two very different worlds. This episode is a rollercoaster ride of heart-stopping action and epic drama, peopled with characters from the beautiful to the bizarre. We meet fish that live on dry land and puffins that must travel 60 miles or more for a single meal, and witness a life-and-death struggle in a technicolour rock pool. In a secluded cove in the Galapagos, sea lions feast on 60kg tuna. It should be impossible - tuna are usually far too fast for sea lions to catch. But here the sea lions club together to herd their prey inshore. Once trapped in the shallows, these huge fish are easy pickings. As the tide recedes in Brazil, lightfoot crabs leap from rock to rock, desperately avoiding the water - their lives depend on it. Moray eels launch themselves from rock pools, jaw gaping. Then octopuses, too. Both crawl across dry rock to set their ambush. Elsewhere, the ever-changing tides create rock pools. But these temporary worlds are a battleground. Predatory starfish turn a magical garden into the stuff of nightmares.

The world of the armoured creatures is equally fascinating and bizarre. Galapagos giant tortoises solve the problem of making love in a suit of armour; a heated sea turtle orgy leaves females in mortal danger and we witness the explosive arrival of a baby pig-nosed turtle. Giant salt-water crocodiles crowd a flooded river-run to fish for mullet and a caiman leads her brood in a touching trek to a communal crèche.

600 miles off the coast of South America in the Pacific Ocean there are 120 tropical islands that erupted from the deep ocean floor more than 10 million years ago. Journey from the lava ramparts to its fiery heart, we'll discover how the Galapagos archipelago became one of the most important areas of biodiversity in the world.
Those swept here by storms and currents survived million to one odds to find untouched islands free from competition. These hardy pioneers evolved in remarkable ways in what's known as Nature's Greatest Experiment. Home to bizarre specialists and unexpected giants, the most precious collection of island creatures on Earth.