On the coast, two worlds collide. Coasts are the most dynamic and challenging habitats in the ocean - that brings great rewards but also great danger. The extraordinary animals that live here must find ingenious ways to cope with two very different worlds. This episode is a rollercoaster ride of heart-stopping action and epic drama, peopled with characters from the beautiful to the bizarre. We meet fish that live on dry land and puffins that must travel 60 miles or more for a single meal, and witness a life-and-death struggle in a technicolour rock pool. In a secluded cove in the Galapagos, sea lions feast on 60kg tuna. It should be impossible - tuna are usually far too fast for sea lions to catch. But here the sea lions club together to herd their prey inshore. Once trapped in the shallows, these huge fish are easy pickings. As the tide recedes in Brazil, lightfoot crabs leap from rock to rock, desperately avoiding the water - their lives depend on it. Moray eels launch themselves from rock pools, jaw gaping. Then octopuses, too. Both crawl across dry rock to set their ambush. Elsewhere, the ever-changing tides create rock pools. But these temporary worlds are a battleground. Predatory starfish turn a magical garden into the stuff of nightmares.

In the second episode, the directors learn their vision for the films is not always permissible by the U.S. government. Wyler is shocked by the racism he encounters against African American soldiers and refused to make a film recruiting black soldiers. Meanwhile, the films' racist depiction of the Japanese versus human depiction of the Germans causes worry for the War Department, which at that time planned to redistribute the Japanese-American population from internment camps into towns across the United States.

The five directors, John Ford, William Wyler, John Huston, Frank Capra, and George Stevens, return to Hollywood after the war but are forever haunted by what they saw. Ford goes on a drinking bender after filming the carnage at D-Day. Stevens is wholly unprepared for the horrors of Dachau and realizes he is not there to film propaganda but to capture evidence of crimes against humanity. Wyler, who lost his hearing during the war, fears his career is over. Huston chronicles soldiers suffering from postraumatic stress disorder in the film Let There Be Light, only to have it suppressed by the U.S. government.

Africa's wildest river is home to the most spectacular wildlife. Hippos fight for territory while herds of elephant, water buffalo and zebra depend on it for life. In the wet season the rains burst the riverbanks and everyone, including people, must move whilst fish swim through the villages. In the dry season the creatures fight over the few pools of water while predators prowl. At its heart it plunges over Victoria Falls and into wild ravines before draining into the Indian Ocean, where storm clouds cycle the water back into the heart of Africa.

Lucy Worsley continues her journey through Russia in the footsteps of the Romanovs, the most powerful royal dynasty in modern European history. In this episode she examines the extraordinary reign of Catherine the Great, and the traumatic conflict with napoleonic France that provides the setting for the novel War and Peace. At the magnificent palace of Peterhof near St Petersburg, Lucy charts the meteoric rise of Catherine the Great, who seized the Russian throne from her husband Peter III in 1762 and became the most powerful woman in the world. Catherine was a woman of huge passions - for art, for her adopted country (she was German by birth) and for her many lovers.
Catherine expanded her empire through military victories overseas, while at home she encouraged education and introduced smallpox inoculation to Russia. But Catherine struggled to introduce deeper reforms, and the institution of serfdom remained largely unchanged. Lucy explains how this injustice fuelled a violent rebellion. Nevertheless, Catherine left Russia more powerful on the world stage than ever. But all she had achieved looked set to be undone when Napoleon invaded Russia in 1812. Lucy relives the pivotal battle of Borodino, when the Russian army finally confronted the French forces; the traumatic destruction of Moscow; and, under Catherine's grandson Alexander, the eventual victory over the French that provided the Romanov dynasty with its most glorious hour.

The years 1825-1918 were bloody and traumatic, a period when four tsars tried - and failed - to deal with the growing pressure for constitutional reform and revolution. In 1861, millions of enslaved serfs were freed by the Tsar-Liberator, Alexander II. But Alexander paid the ultimate penalty for opening the Pandora's box of reform when he was later blown up by terrorists on the streets of St Petersburg. Elsewhere, there was repression, denial, war and - in the case of the last tsar, Nicholas II - a fatalistic belief in the power of God, with Nicholas's faith in the notorious holy man Rasputin being a major part in his undoing. Lucy Worsley also details the chilling murder of Nicholas and his family in 1918, and asks whether all of this horror have been avoided.