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The world's deserts are lands of extremes that force animals to come up with ingenious ways of coping with hostile conditions, giving rise to the most incredible survival stories on earth. A pride of desert lions are so hungry they risk hunting a giraffe several times their size, while male sandgrouse fly 120 miles each day to the nearest waterhole and dice with death to collect water for their chicks. Filmed for the first time, a tiny bat does battle with one of the world's deadliest scorpions, and in Madagascar, a locust swarm of biblical proportions is seen as never before.

Grasslands cover one quarter of all land and support the vast gatherings of wildlife, but to survive here animals must endure the most hostile seasonal changes on the planet. From Asia's bizarre-looking Saiga antelope to the giant anteaters of Brazil, grassland animals have adapted in extraordinary ways to cope with these extremes. In the flooded Okavango, lions take on formidable buffalo in epic battles, on the savannah bee-eaters take advantage of elephants to help catch insects and, on the freezing northern tundra, caribou embark on great migrations shadowed by hungry Arctic wolves.

Cities are growing at a faster rate than any other habitat on Earth. They may seem an unlikely place for animals to thrive, but they can be a world of surprising opportunity. Leopards prowl the streets of Mumbai, peregrine falcons hunt amongst New York's skyscrapers, and a million starlings perform spectacular aerial dances over Rome. In Jodhpur, langurs are revered as religious deities and in Harar, locals live in harmony with wild hyenas. Many animals, however, struggle to cope in the urban jungle. As the architects of this environment, can humans choose to build cities that are homes for both them and wildlife?

Nicholas felt enormous pressure to continue the dynasty and expand the empire. He saw his opportunity in the Far East. This brought him into conflict with a rising power, Japan. Russia's disastrous war against Japan prompts civil unrest. The birth of Prince Alexei is incredibly important. He's the long-awaited son that they had hoped so much for. But when Alexei is diagnosed with haemophilia, Siberian mystic Rasputin steps in. He's very good at realizing when something is open that will enable him to increase his power and wealth. The pressure of keeping the secret of Alexei's illness affects the political decisions that Nicky and Alix made for the rest of their reign.

The Romanovs retreat into a bubble of luxury while the czar's forces brutally suppress protests by a starving populace. Rasputin's influence grows, never mind the rumors and scandals. Many people believe Prime minister Peter Stolypin is the last hope of the Romanov Dynasty as reformer-in-chief. He is horrified by the rise of Rasputin. The fight between Stolypin and Rasputin is a battle for influence over Nicholas II. But Stolypin is shot in the Kiev Opera in 1911.

In 1914, Russia entered World War I in response to Austria-Hungary's declaration of war on Russia's ally Serbia. This was fought across multiple fronts while isolated from its allies. The Battle of Tannenberg, where the entire Russian Second Army was annihilated, cast an ominous shadow over the empire's future. The already-existing public distrust of the regime was deepened by the rising costs of the war and rumours of corruption. World War I takes Nicholas to the front. Back home, outrage grows over high casualties and Alexandra and Rasputin's political intrigues.