Intelligence and adaptability allow primates to tackle the many challenges of life, and this is what makes our closest relatives so successful. This resourcefulness has enabled primates to conquer an incredible diversity of habitat. Hamadryas baboons live on the open plains of Ethiopia in groups up to 400 strong. Strength in numbers gives them some protection from potential predators. But, should their path cross with other baboon troops, it can lead to all-out battle, as males try to steal females from one another, and even settle old scores. Japanese macaques are the most northerly-dwelling primates and they experience completely different challenges. Some beat the freezing conditions by having access to a thermal spa in the middle of winter. But this privilege is only for those born of the right female bloodline. For western lowland gorillas, it's the male silverback that leads his family group in the rich forests of the Congo basin. He advertises his status to all with a powerful chest-beating display. Most primates are forest dwellers, and one of the strangest is the tarsier – the only purely carnivorous primate. As it hunts for insects the tarsier leaps from tree to tree in the dead of night, using its huge forward-facing eyes to safely judge each jump. Good communication is essential for success in primate society.

Sagan discusses the story of the Heike crab and artificial selection of crabs resembling samurai warriors, as an opening into a larger discussion of evolution through natural selection (and the pitfalls of intelligent design). Among the topics are the development of life on the Cosmic Calendar and the Cambrian explosion; the function of DNA in growth; genetic replication, repairs, and mutation; the common biochemistry of terrestrial organisms; the creation of the molecules of life in the Miller-Urey experiment; and speculation on alien life (such as life in Jupiter's clouds). In the Cosmos Update ten years later, Sagan remarks on RNA also controlling chemical reactions and reproducing itself and the different roles of comets (potentially carrying organic molecules or causing the Cretaceous–Paleogene extinction event).

The theory of Némesis, a star that orbits the Sun and causes catastrophic events is explored. Nemesis is a hypothetical red dwarf or brown dwarf, originally postulated in 1984 to be orbiting the Sun at a distance of about 95,000 AU (1.5 light-years) somewhat beyond the Oort cloud, to explain a perceived cycle of mass extinctions in the geological record, which seem to occur more often at intervals of 26 million years.

The contests between predators and prey are the most dramatic events in nature. Taking an intimate and detailed look at the remarkable strategies employed by hunters to catch their prey and the hunted to escape. Sir David Attenborough narrates. "The Hardest Challenge" reveals the extraordinary range of techniques predators use to catch their prey - from a leopard using all its powers of stealth to stalk impala in broad daylight to wild dogs, whose tactic is to wear down their prey over long distances; from Nile crocodiles, the planet's most patient predators, to killer whales who use teamwork and intelligence to take on humpback whales. But even with these finely tuned strategies, the outcome is far from certain. Surprisingly, most predators fail most of the time.

We see how the Germans they lived, fought and worshippped their gods. Intricate 3d animation shows how they built their settlements, buried their kings, vanquished their enemies.

The Dutch Golden Age saw map-making reach a fever pitch of creative and commercial ambition. This was the era of the first ever Atlases - elaborate, lavish and beautiful. This was the great age of discovery and marked an unprecedented opportunity for mapmakers who sought to record and categorise the newly acquired knowledge of the world. Rising above the many mapmakers in this period was Gerard Mercator, inventor of the Mercator projection, who changed mapmaking forever when he published his collection of world maps in 1598 and coined the term 'Atlas'. The programme looks at some of the largest and most elaborate maps ever produced, from the vast maps on the floor of the Royal Palace in Amsterdam, to the 24 volume atlas covering just the Netherlands, to the largest Atlas in the world, The Klencke Atlas. It was made for Charles II to mark his restoration in 1660. But whilst being one of the British Libraries most important items, it is also one of its most fragile so hardly ever opened. This is a unique opportunity to see inside this enormous and lavish work, and see the world through the eyes of a King.