The Arctic is the harshest environment on Earth: little food grows, it's dark for months on end, and temperatures stay well below freezing for much of the year. Yet four million people manage to survive here. Human Planet tells remarkable stories of extraordinary people who make their homes in nature's deep freeze. In springtime, Amos and Karl-Frederik set out across the sea ice with their dogs to catch a real-life sea monster: a Greenland shark! Inuit mussel-gatherers venture underneath the sea ice at low tide for a perilous race against time as they gather their food. And the children of Churchill, Manitoba, set out on the most dangerous trick or treating Halloween in the world: they risk coming face-to-face with deadly polar bears on the streets of their town. Who'll get the tastiest snack?

Grasslands feed the world. Over thousands of years, we humans have learned to grow grains on the grasslands and domesticate the creatures that live there. Our success has propelled our population to almost seven billion people. But this episode reveals that, even today, life in the 'Garden of Eden' isn't always rosy. We walk with the Dorobo people of Kenya as they bravely attempt to scare off a pride of hungry lions from their freshly caught kill. We gallop across the Steppe with extraordinary Mongolian horsemen who were 'born in the saddle'. And in a perfect partnership with nature built up over generations Maasai children must literally talk to the birds! The honeyguide leads them to find sweet treats, but they'll have to repay the favour.

Sam Neill explores humanity's long-term future - a future where we make new homes for ourselves on other worlds as we flee from the slow but inevitable death of our sun.

Are the wealthy just born in the right place at the right time? Are the poor victims of a system designed to keep them down? Or do physics and biology determine who is rich and who is poor? Throughout history, distribution of wealth is governed by hidden forces: DNA, environmental stress, patterns of human migration and even the laws of thermodynamics! Nature seems to demand winners and losers in life. But does this mean greed is king, and the rich can take what they want? Or is cooperation – between microorganisms, monkeys, and humans – more essential to survival of a species?

In the final challenge of the game of life, raising offspring is the ultimate prize. Continuing the line through the next generation, is the next best thing to immortality. But it’s far from easy... Some parents must risk their own lives for their offspring.

In this final episode we complete our journey, travelling back from the March equinox to the end of June. Kate Humble is in the Arctic at a place where spring arrives with a bang, whilst Helen Czerski chases a tornado to show how the earth's angle of tilt creates the most extreme weather on earth.