On the coast, two worlds collide. Coasts are the most dynamic and challenging habitats in the ocean - that brings great rewards but also great danger. The extraordinary animals that live here must find ingenious ways to cope with two very different worlds. This episode is a rollercoaster ride of heart-stopping action and epic drama, peopled with characters from the beautiful to the bizarre. We meet fish that live on dry land and puffins that must travel 60 miles or more for a single meal, and witness a life-and-death struggle in a technicolour rock pool. In a secluded cove in the Galapagos, sea lions feast on 60kg tuna. It should be impossible - tuna are usually far too fast for sea lions to catch. But here the sea lions club together to herd their prey inshore. Once trapped in the shallows, these huge fish are easy pickings. As the tide recedes in Brazil, lightfoot crabs leap from rock to rock, desperately avoiding the water - their lives depend on it. Moray eels launch themselves from rock pools, jaw gaping. Then octopuses, too. Both crawl across dry rock to set their ambush. Elsewhere, the ever-changing tides create rock pools. But these temporary worlds are a battleground. Predatory starfish turn a magical garden into the stuff of nightmares.

Lucy Worsley continues her journey through Russia in the footsteps of the Romanovs, the most powerful royal dynasty in modern European history. In this episode she examines the extraordinary reign of Catherine the Great, and the traumatic conflict with napoleonic France that provides the setting for the novel War and Peace. At the magnificent palace of Peterhof near St Petersburg, Lucy charts the meteoric rise of Catherine the Great, who seized the Russian throne from her husband Peter III in 1762 and became the most powerful woman in the world. Catherine was a woman of huge passions - for art, for her adopted country (she was German by birth) and for her many lovers.
Catherine expanded her empire through military victories overseas, while at home she encouraged education and introduced smallpox inoculation to Russia. But Catherine struggled to introduce deeper reforms, and the institution of serfdom remained largely unchanged. Lucy explains how this injustice fuelled a violent rebellion. Nevertheless, Catherine left Russia more powerful on the world stage than ever. But all she had achieved looked set to be undone when Napoleon invaded Russia in 1812. Lucy relives the pivotal battle of Borodino, when the Russian army finally confronted the French forces; the traumatic destruction of Moscow; and, under Catherine's grandson Alexander, the eventual victory over the French that provided the Romanov dynasty with its most glorious hour.

The years 1825-1918 were bloody and traumatic, a period when four tsars tried - and failed - to deal with the growing pressure for constitutional reform and revolution. In 1861, millions of enslaved serfs were freed by the Tsar-Liberator, Alexander II. But Alexander paid the ultimate penalty for opening the Pandora's box of reform when he was later blown up by terrorists on the streets of St Petersburg. Elsewhere, there was repression, denial, war and - in the case of the last tsar, Nicholas II - a fatalistic belief in the power of God, with Nicholas's faith in the notorious holy man Rasputin being a major part in his undoing. Lucy Worsley also details the chilling murder of Nicholas and his family in 1918, and asks whether all of this horror have been avoided.

Dr Janina Ramirez shows how medieval manuscripts gave power to the king and united the kingdom in an age of plague, warfare and rebellion, discovers that Edward III used the manuscripts he read as a boy to prepare him for his great victory at the battle of Crecy and reveals how a vigorous new national identity bloomed during the 100 Years War with France. In the British Library's Royal Manuscripts collection Dr Ramirez finds out that magnificent manuscripts like the Bedford Hours, taken as war booty from the French royal family, were adapted for the education of English princes. She also explores how knowledge spread through a new form of book - the encyclopaedia.

Every year grizzly bear families in North America depend for their survival on a spectacular natural event: the return of hundreds of millions of salmon from the Pacific Ocean to the mountain streams where they were born. The salmon travel thousands of miles to spawn and then die. The great run not only provides food for bears, but for killer whales, wolves, bald eagles, and even the forest itself. The question is: will the salmon return in time to keep hungry bears alive?
A mother grizzly and her cubs emerge from their den high in snowy Alaskan mountains. Filming from the air the team capture a TV first, following the bears as they negotiate a near vertical slope on their journey to the coast where they await the return of the salmon. Meanwhile, the salmon are making their way to the to river mouths where they must swim upstream and against the current. The programme reveals how they tackle the torrents and leap over waterfalls, a feat equivalent to a human jumping over a house. Dozens of hungry bears eagerly await the salmon that make it up river. In another TV first, underwater cameras record the ingenuity and fancy footwork they use to collect dead salmon from the bottom of deep pools.

Each year more than one million wildebeest and zebra invade the Serengeti grasslands, making it a paradise for the predators that live there. But what happens when the herds move off again? We follow the moving story of one lion family's struggle to survive until the return of the great migration. Nature's Great Events tells the story of the epic trek of herds that follow the rains to fresh pastures, and the tale of the predators they leave behind.
The crew captures the desperate plight of a single pride of lions, revealing a different side to the Serengeti. Rather than being a predators' paradise, it is a land in constant change, with wildebeest following the rains and leaving the lions to tough it out. The Ntudu pride has seven cubs, and is already suffering as the wildebeest leave to find fresh pastures. The four pride females struggle to find enough food for their hungry offspring. As weeks turn to months, the pride members become more emaciated and frailer, and the number of cubs dwindles to just two. As the herds begin to return, the plains reveal one final secret. For the first time since 1967 the Serengeti's only active volcano, Ol Doinyo Lengai, begins to billow ash and smoke. Filmed from the air, the team captures the exciting action. Fertilised by the volcanic ash over millions of years, these short grass plains are among the most productive grasslands in the world. After months of hardship, the pride's tragic story, through sickness, drought and fire, is over as the herds return, providing plentiful food.