Series concludes with the focus shifting to the United States in the post-war years of the 1950s and beyond. Beginning with arguably the most notorious work of 20th century classical music, John Cage's 'silent' composition 4'33", it looks at how a series of maverick Americans re-invented the sound of classical music into a more simple form, bringing back harmonies and rhythms that made it increasingly popular with audiences across the world. It also examines how this music found its way into a spiritual realm, with the strain of pared-down religious composition that came to be known as 'holy minimalism'. From the Maverick concert hall in Woodstock, New York to an Orthodox cathedral in Estonia to a car park in Peckham, south London, the story is told by a stellar line-up of contributors including Philip Glass, Steve Reich, John Adams, Arvo Pärt and John Tavener.

For the animals in the polar regions, autumn means dramatic battles and epic journeys. Time is running out - the Arctic Ocean is freezing over and the sea ice is advancing at 2.5 miles per day around Antarctica. Polar bears gather in large numbers on the Arctic coast as they wait for the return of the ice. Soon, tempers fray and violent sparring contests break out. Meanwhile 2,000 beluga whales head for one special estuary, a gigantic 'whale spa' where they will thrash their snow-white bodies against the gravel and exfoliate. Inland, the tundra undergoes a dramatic transformation from green to fiery red. Here, musk ox males slam head-first into each other with the force of a 30mph car crash as they struggle to defend their harems. Frisky young caribou males play a game of 'grandma's footsteps' as they try to steal the boss's female. Down in Antarctica, Adelie penguin chicks huddle together in creches. When a parent returns from fishing, it leads its twins on a comical steeplechase - sadly there's only enough for one, so the winner gets the meal. Two months later and the chicks are fully feathered apart from downy Mohican hairdos - they're ready to take their first swim - reluctantly though, as it seems penguins are not born with a love of water! And with good reason - a leopard seal explodes from the sea and pulls one from an ice floe, a hunting manoeuvre that has never been filmed before. As winter approaches and everyone has left, the giant emperor penguin arrives and makes an epic trek inland to breed. The mothers soon return to the sea leaving the fathers to hold the eggs and endure the coldest winter on earth.

In late Cretaceous Canada, in what will be known as Dinosaur Provincial Park, a Daspletosaurus stalks a Chasmosaurus in a forest, but loses the element of surprise and is forced to retreat. The Chasmosaurus comes across a younger Daspletosaurus, before being ambushed by a group of the tyrannosaurids. The episode then cuts to the high Arctic, where Edmontosaurus are hunted by a large species of Troodon. The theropods attack at night, separating a juvenile from the herd and severely wounding it, only to be driven away by an adult. In the morning, they return to eat the carcass of the juvenile, which died during the night. The episode returns to the Daspletosaurus, who chase and bring down the Chasmosaurus. The larger adults bully the youngsters off the carcass, forcing them to wait until they have finished. The episode then cuts to Madagascar, where a mother Majungasaurus (an Abelisaurid) and her two offspring chase a group of Rahonavis off a carcass. However, they are temporarily driven off themselves by a male Majungasaurus. But, after he steals some food from one of the young, the female attacks him, before she and her young cannibalise his body. The episode returns once again to North America, where the Daspletosaurus are waiting for the annual migration of Centrosaurus. They attack during a rainstorm, killing some of the ceratopsians. The Centrosaurus make it to a flooded river and begin to swim across, and although many make it to the other side, some are caught by giant crocodilians or are severely wounded by floating debris and thus drown, or drown for unseen reasons. In the morning, the carcasses attract scavengers, including the Daspletosaurus. A montage is then shown of Daspletosaurus and Majungasaurus, the narrator saying that together, the tyrannosaurids and abelisaurids were the last of the killer dinosaurs.

The final episode explores dinosaurs' extraordinary ability to survive. Featured dinosaurs include the bizarre magyarosaurus, which lived in the shadow of the biggest flying animal - hatzegopteryx - and showed an amazing adaptation to island life; and the weird nothronychus, a carnivore that gave up meat eating. This astonishing capacity to evolve into ever more diverse and bizarre forms meant that dinosaurs not only spread throughout the world, but also dominated life upon it for more than 160 million years. It was only an unprecedented extraterrestrial event that finally saw the end of planet dinosaur.

During 1953-1958, as the Cold War intensifies, Korolev is asked to build a rocket capable of carrying a five-ton warhead to America - he designs and constructs the R-7 Semyorka, and is later allowed to use it to launch the first satellite, Sputnik 1. Meanwhile, von Braun struggles to persuade the US government to allow him to launch his own satellite - after Sputnik's launch and the failure of the US Navy to launch a Vanguard satellite, he is finally allowed to launch the first American satellite, Explorer 1.

The rainforest is home to more species of plants and animals than any other habitat on the planet. But for humans, life there is not as easy as it looks. Life in the trees requires great skill, ingenuity and sheer bravery. The Matis of Brazil carve 4-metre-long blow-pipes to hunt monkeys - in near total silence. Deep in the Congo forests, Tete defies death by scaling a giant tree using nothing more than a liana vine, and he must then negotiate an angry swarm of bees - all to collect honey for his family. Three children from Venezuela's Piaroa tribe venture deep into the jungle to hunt tarantulas - to toast for lunch! In West Papua the Korowai tribe show-off their engineering skills by building a high-rise home 35 metres up in the tree tops. Most memorable of all, in Brazil we join a unique monitoring flight in search an un-contacted tribe...