The contests between predators and prey are the most dramatic events in nature. Taking an intimate and detailed look at the remarkable strategies employed by hunters to catch their prey and the hunted to escape. Sir David Attenborough narrates. "The Hardest Challenge" reveals the extraordinary range of techniques predators use to catch their prey - from a leopard using all its powers of stealth to stalk impala in broad daylight to wild dogs, whose tactic is to wear down their prey over long distances; from Nile crocodiles, the planet's most patient predators, to killer whales who use teamwork and intelligence to take on humpback whales. But even with these finely tuned strategies, the outcome is far from certain. Surprisingly, most predators fail most of the time.

After filming for three years, Planet Earth finally captures the shy Mongolian gazelle. Only a handful of people have witnessed its annual migration. Don't miss the bizarre-looking Tibetan fox, captured on film for the first time. Over six weeks the team follow a pride of 30 lions as they attempt to hunt elephants. Using the latest night vision equipment, the crew film the chaotic battles that ensue at close quarters.

A look at the solar system's largest planet, Jupiter; its formation and composition and its mini-solar system of over 60 moons – some of which may have the potential to support extra-terrestrial life.

Silk is the invertebrates' great invention, used in a range of ways from from the protective stalks of lacewing eggs to the amazing hanging threads of New Zealand's 'glow worms'. Spiders, though, have taken silk-spinning to extremes. The common wolf spider has no web, but the female is a gentle parent, encasing her eggs in silk and carrying the precious bundle wherever she goes. The bolas spider uses a ball of sticky silk soaked in a copy of moth pheromone to lure its prey. Millions of communal spiders live and feed together in a vast, towering web - an arachnophobe's nightmare.

Ideas about time and space are explored in the changes that constellations undergo over time, the redshift and blue shift measured in interstellar objects, time dilation in Albert Einstein's theory of relativity, the designs of both Leonardo da Vinci and spacecraft that could travel near light speed, time travel and its hypothetical effects on human history, the origins of the Solar System, the history of life, and the immensity of space. In Cosmos Update, the idea of faster-than-light travel by wormholes (researched by Kip Thorne and shown in Sagan’s novel Contact) is discussed.

Sagan reflects on the future of humanity and the question of 'who speaks for Earth?' when meeting extraterrestrials. He discusses the very different meetings of the Tlingit people and explorer Jean-François de La Perouse with the destruction of the Aztecs by Spanish conquistadors, the looming threat of nuclear warfare, and the threats shown by destruction of the Library of Alexandria and the murder of Hypatia. The episode ends with an overview of the beginning of the universe, the evolution of life, and the accomplishments of humanity and makes a plea to mankind to cherish life and continue its journey in the cosmos. The Cosmos Update notes the preliminary reconnaissance of planets with spacecraft, the fall of the Berlin Wall and the end of apartheid in South Africa, and measures towards the reduction of nuclear weapons.