In the last episode of the series, lavish imperial games at the Colosseum are highlighted by the emperor's bloody victories, but his paranoid abuses of power lead to a final betrayal.

In the second episode, Andrew Carnegie builds an empire around steel, but finds himself struggling to save face after the ruthless tactics of his business partner, Henry Frick, result in both the Johnstown Flood as well as the bloody 1892 strike at the Homestead Steel Works.

In the fourth part of the series, Diarmaid MacCulloch makes sense of the Reformation, and of how a faith based on obedience and authority gave birth to one based on individual conscience. He shows how Martin Luther wrote hymns to teach people the message of the Bible, and how a tasty sausage became the rallying cry for Swiss Reformer Ulrich Zwingli to tear down statues of saints, allow married clergy and deny that communion bread and wine were the body and blood of Christ.

The years 1825-1918 were bloody and traumatic, a period when four tsars tried - and failed - to deal with the growing pressure for constitutional reform and revolution. In 1861, millions of enslaved serfs were freed by the Tsar-Liberator, Alexander II. But Alexander paid the ultimate penalty for opening the Pandora's box of reform when he was later blown up by terrorists on the streets of St Petersburg. Elsewhere, there was repression, denial, war and - in the case of the last tsar, Nicholas II - a fatalistic belief in the power of God, with Nicholas's faith in the notorious holy man Rasputin being a major part in his undoing. Lucy Worsley also details the chilling murder of Nicholas and his family in 1918, and asks whether all of this horror have been avoided.

From steamy jungles to dry deserts, amphibians have taken their first footsteps onto land using their bizarre life histories to break their ties with the water and invade the earth.
This episode delves into the extraordinary and intimate lives of the soft skinned amphibians. This includes the marsupial frogs, a peculiar species where the father carries his young in pouches and then gives birth. It also features warring giant salamanders over a metre long and show-off newts that offer displays just like underwater birds of paradise.

David Attenborough reveals the fascinating lives of snakes, the most misunderstood group of reptiles. A CCTV stakeout of wild rattlesnakes hunting shows, for the first time, what sophisticated predators they truly are. Attenborough is also on the receiving end of a spitting cobra's chemical weapons system, while the surprisingly beautiful and tender side of snakes is displayed in the sinuous courtship of kingsnakes, and the water birth of 15 beautiful yellow anaconda babies.