David Attenborough concludes his epic history of the evolution of flight with an exploration of the highly advanced fliers that dominate our skies today – the extraordinarily diverse skills of the birds, and the sonar-guided precision of the bats. He encounters some of our planet’s most remarkable fliers – from peregrine falcons dive-bombing starlings over Rome, to hummingbirds hovering in the cloud forests of Ecuador. In a spectacular finale, he visits Gomantong cave in Borneo to witness the mass exodus of a million bats.

Moving on to the Late Eocene period 36 million years ago and mammals have prospered and are now the largest creatures on land and sea. This is an era of animals like andrewsarchus, the biggest mammal carnivore ever to walk on land, and the brontotheres, small-brained herbivores. It is in the sea, however, that the most monstrous mammals of all can be found. We follow the fate of a female basilosaurus, a huge serpent-like early whale, but nothing like the gentle filter feeding whales of the 21st century. Four times the length of the great white shark, with jaws to match, she is every inch a killer.

25 million years ago the biggest land mammals of all time, the indricotheres, stalked the Earth. Up to seven metres tall and weighing 15 tonnes, adults were too big to be eaten by any predator of the time. Only in the first few years of its life was indricotheres vulnerable. The programme follows the fate of a calf from his traumatic birth to see whether it can survive droughts, killer hogs and hyaenodon - a predator the size of a rhino with jaws that could crush a rock.

Moving on to Ethiopia 3.2 million years ago, we witness the beginnings of mankind via a group of australopithecus - a type of ape which, like us, walks upright on two legs. But unlike us, these early members of the human family weren't predators, they were prey. Things get worse for the group as they are hunted by a sabre-tooth cat called dinofelis and fall victim to other dangers such as malaria, rival australopithecus and a rampaging 14-tonne deinotherium.

Back one million years ago, South America was a continent of exotic oversized creatures found nowhere else on Earth - huge terror birds, giant ground sloths and spiky-tailed relatives of the armadillo as big as cars. The deadliest animal of them all was smilodon, the largest of the sabre-tooth cats, with canines like carving knives. The programme follows the fortunes of an individual male, Half Tooth. Ousted from his clan by a pair of rival males, his life suddenly becomes a struggle to survive in this alien world.

Diarmaid MacCulloch's own life story makes him a symbol of a distinctive feature about Western Christianity - scepticism, a tendency to doubt which has transformed both Western culture and Christianity. In the final programme in the series, he asks where that change came from. He challenges the simplistic notion that faith in Christianity has steadily ebbed away before the relentless advance of science, reason and progress, and shows instead how the tide of faith perversely flows back in. Despite the attacks of Newton, Voltaire, the French Revolutionaries and Darwin, Christianity has shown a remarkable resilience. The greatest damage to Christianity was actually inflicted to its moral credibility by the two great wars of the 20th century and by its entanglement with Fascism and Nazism. And yet it is during crisis that the Church has rediscovered deep and enduring truths about itself, which may even be a clue to its future.