The second episode looks at the challenges of hunting in the Arctic, the most seasonal place on Earth. To a predator, seasonal change is a problem. It means that all the parameters of the hunt - the conditions, the strategies, the prey - change too. The only option for the Arctic's top predators, the wolf, the Arctic fox and the polar bear, is to continually adapt to their changing world, exploiting the good times and enduring the bad.

25 million years ago the biggest land mammals of all time, the indricotheres, stalked the Earth. Up to seven metres tall and weighing 15 tonnes, adults were too big to be eaten by any predator of the time. Only in the first few years of its life was indricotheres vulnerable. The programme follows the fate of a calf from his traumatic birth to see whether it can survive droughts, killer hogs and hyaenodon - a predator the size of a rhino with jaws that could crush a rock.

In the Land of the Cave Bear, Alice ventures to the parts of the northern hemisphere, hit hardest by the cold - Europe and Siberia. High in the mountains of Transylvania, a cave sealed for thousands of years reveals grisly evidence for a fight to the death between two staving giants, a cave bear and a cave lion. Yet Alice discovers that for woolly rhinos and woolly mammoths, the Ice Age created a bounty. The Mammoth Steppe, a vast tract of land which went half way round the world, provided food all year round, for those that liked the cold. It was these mammoths that Europe's most dangerous predators hunted for their survival.

Marine invertebrates are some of the most bizarre and beautiful animals on the planet, and thrive in the toughest parts of the oceans. Divers swim into a shoal of predatory Humboldt squid as they emerge from the ocean depths to hunt in packs. When cuttlefish gather to mate, their bodies flash in stroboscopic colours. Time-lapse photography reveals thousands of starfish gathering under the Arctic ice to devour a seal carcass. A giant octopus commits suicide for her young. A camera follows her into a cave which she walls up, then she protects her eggs until she starves. The greatest living structures on earth, coral reefs, are created by tiny animals in some of the world's most inhospitable waters.

Mammals' ability to learn new tricks is the key to survival in the knife-edge world of hunters and hunted. In a TV first, a killer whale off the Falklands does something unique: it sneaks into a pool where elephant seal pups learn to swim and snatches them, saving itself the trouble of hunting in the open sea. Slow-motion cameras reveal the star-nosed mole's newly-discovered technique for smelling prey underwater: it exhales then inhales a bubble of air ten times per second. Young ibex soon learn the only way to escape a fox - run up an almost vertical cliff face - and young stoats fight mock battles, learning the skills that make them one of the world's most efficient predators.

Intelligence and adaptability allow primates to tackle the many challenges of life, and this is what makes our closest relatives so successful. This resourcefulness has enabled primates to conquer an incredible diversity of habitat. Hamadryas baboons live on the open plains of Ethiopia in groups up to 400 strong. Strength in numbers gives them some protection from potential predators. But, should their path cross with other baboon troops, it can lead to all-out battle, as males try to steal females from one another, and even settle old scores. Japanese macaques are the most northerly-dwelling primates and they experience completely different challenges. Some beat the freezing conditions by having access to a thermal spa in the middle of winter. But this privilege is only for those born of the right female bloodline. For western lowland gorillas, it's the male silverback that leads his family group in the rich forests of the Congo basin. He advertises his status to all with a powerful chest-beating display. Most primates are forest dwellers, and one of the strangest is the tarsier – the only purely carnivorous primate. As it hunts for insects the tarsier leaps from tree to tree in the dead of night, using its huge forward-facing eyes to safely judge each jump. Good communication is essential for success in primate society.